Utente:Esculapio/Sandbox/Melolonthinae

Melolonthinae MacLeay, 1819 ^[1]

Tribù Ablaberini Burmeister, 1855 [=Camentini Machatschtke, 1959]
- Ablabera Erichson, 1848 [Dejean 1833?] [=Parablabera Peringuey, 1904] [new?]
  - Ablabera splendida (Fabricius, 1781) [present designation; BMNH] [=Melolontha splendida Fabricius, 1781]
  - Ablabera transvaalica Peringuey, 1904 [present designation; SAMC]
- Ablaberoides Blanchard, 1850
  - Ablaberoides aeneus Blanchard, 1850 [by monotypy; MNHN]
- Achelyna Erichson, 1847
  - Achelyna clypeata Burmeister, 1855 [by subsequent monotypy; Burmeister 1855; MLUH] [=Arraphytarsa Peringuey, 1904] [Arrow 1925]
  - Allokotarsa damarina (Peringuey, 1904) [by monotypy; SAMC] [=Arraphytarsa damarina Peringuey, 1904]
  - Allokotarsa rotundicollis Peringuey, 1904 [present designation; SAMC]
  - Allokotarsa straminea (Peringuey, 1904) [present designation; SAMC] [=Alogistotarsa straminea Peringuey, 1904]
- Apocamenta Brenske, 1903
  - Apocamenta variolosa (Brenske, 1897) [present designation; ZMHB] [=Camenta variolosa Brenske, 1897]
- Archocamenta Brenske, 1895
  - Archocamenta flava Brenske, 1895 [by monotypy; ZMHB]
- Brachymys Thomson, 1858 [=Apocamenta Brenske, 1903] [Burgeon 1945]
  - Brachymys cameruna (Brenske, 1903) [present designation; ZMHB] [=Apocamenta cameruna Brenske, 1903]
  - Brachymys pubens Thomson, 1858 [by monotypy; ??]
- Camenta Erichson, 1848
  - Camenta obesa Burmeister, 1855 [present designation; MLUH]
- Cyrtocamenta Brenske, 1897
  - Cyrtocamenta flavescens Brenske, 1897 [present designation; ZMHB]
- Diplotropis Boheman, 1857
  - Diplotropis nigrina Fahraeus, 1857 [present designation; NHRS]
- Empecamenta Brenske, 1895 [?; as subgenus of Camenta Peringuey 1904; Machatschke 1959]
  - Empecamenta pilifera Brenske, 1895 [by monotypy; ZMHB]
- Eucamenta Peringuey, 1904 [as subgenus of Camenta Peringuey 1904; Machatschke 1959]
  - Eucamenta castanea (Boheman, 1857) [present designation; SAMC] [=Ablabera castanea Boheman, 1857]
- Gamka Peringuey, 1904
  - Gamka minuta Peringuey, 1904 [by monotypy; SAMC]
- Gryphonycha Peringuey, 1904
  - Gryphonycha puberula (Fahraeus, 1857) [by monotypy; NHRS] [=Serica puberula Fahraeus, 1857]
- Hybocamenta Brenske, 1898 [as subgenus of Camenta Peringuey 1904; Machatschke 1959] [=Liocamenta Kolbe, 1914] [Moser 1919]
  - Hybocamenta benitoana Brenske, 1898 [present designation; ZMHB]
  - Hybocamenta consentanea (Kolbe, 1914) [by monotypy; ZMHB] [=Liocamenta consentanea Kolbe, 1914]
- Idaecamenta Peringuey, 1904
  - Idaecamenta jucunda Peringuey, 1904 [by monotypy; SAMC]
- Idaeserica Peringuey, 1904
  - Idaeserica gratula Peringuey, 1904 [by monotypy; SAMC]
- Isocamenta Kolbe, 1914
  - Isocamenta schubotzi Kolbe, 1914 [by subsequent designation; Zool. Rec.1968; ZMHB]
- Kadlecia Frey, 1965b
  - Kadlecia heptaphylla Frey, 1965 [by monotypy; ZSMC]
- Leribe Peringuey, 1904
  - Leribe vesca Peringuey, 1904 [by monotypy; SAMC]
- Makalaka Peringuey, 1908
  - Makalaka rhodesiana Peringuey, 1908 [by monotypy; SAMC]
- Melanocamenta Brenske, 1899
  - Melanocamenta bomuana Brenske, 1899 [by monotypy; ZMHB]
- Microcamenta Brenske, 1898
  - Microcamenta transvaalensis Brenske, 1898 [by monotypy; ZMHB]
- Mitracamenta Brenske, 1903
  - Mitracamenta lineella Brenske, 1903 [by monotypy; ZMHB]
- Neocamenta Burgeon, 1945
  - Neocamenta flaviceps Burgeon, 1945 [by monotypy; MRAC]
- Oocamenta Peringuey, 1904

SUBGENUS Oocamenta Peringuey, 1904

- - Oocamenta rufiventris (Burmeister, 1855) [by monotypy; SAMC] [=Camenta rufiventris Burmeister, 1855]

SUBGENUS Pseudocamenta Peringuey, 1904

- - Oocamenta transvaalica Peringuey, 1904 [by monotypy; SAMC]
- Pachycamenta Brenske, 1895
  - Pachycamenta gestro Brenske, 1895 [by monotypy; ZMHB]
- Pachychilecamenta Brenske, 1903
  - Pachychilecamenta rufa Brenske, 1903 [by monotypy; ZMHB]
- Paracamenta Peringuey, 1904
  - Paracamenta bohemani (Brenske, 1897) [present designation; ZMHB][=Camenta bohemani Brenske, 1897]
- Pentecamenta Brenske, 1897
  - Pentecamenta salaama Brenske, 1897 [present designation; ZMHB]
- Pericamenta Peringuey, 1904
  - Pericamenta paupercula Peringuey, 1904 [by monotypy; SAMC]
- Phyllocamenta Moser, 1918a
  - Phyllocamenta pilosa (Quedenfeldt, 1884) [by monotypy; ZMHB] [=Camenta pilosa Quedenfeldt, 1884]
- Synacta Fairmaire, 1899
  - Synacta corrugata Fairmaire, 1899 [by monotypy; MNHN]
- Tarsocamenta Moser, 1924
  - Tarsocamenta kapiriana Moser, 1924 [by monotypy; ZMHB]
- Tulbaghia Peringuey, 1904
  - Tulbaghia lighfooti Peringuey, 1904 [present designation; SAMC]
Tribù Alvarengiini Frey, 1975a
- Alvarengius Frey, 1975a
  - Alvarengius silphodes Frey, 1975a [by monotypy; ZSMC?]
Tribù Automoliini Britton, 1978 [key to genera Britton 1957]
- Aplopsis Blanchard, 1850 [LR Britton 1957] [=Atermonocheila Blackburn, 1907] [Britton 1987][=Colpocrania Burmeister, 1855] [Britton 1987] [=Haplopsis Blanchard, 1850] [unjustified emendation Gemminger & Harold 1869] [=Ocnodus Burmeister, 1855] [Britton 1987]
  - Aplopsis decipiens (Burmeister, 1855) [by monotypy; MLUH?] [=Ocnodus decipiens Burmeister, 1855]
  - Aplopsis lineoligera (Blanchard, 1850) [by monotypy; MNHN] [=Haplopsis lineoligera Blanchard, 1850]
  - Aplopsis longipes (Blackburn, 1907) [by monotypy; SAMA] [=Atermonocheila longipes Blackburn, 1907]
  - Aplopsis pilosa (Burmeister, 1855) [here designated; MLUH] [=Colpocrania pilosa Burmeister, 1855]
- Automolius Britton, 1978 [LR Britton 1957; n.n. pro Automolus] [=Automolus Burmeister, 1855] [not Reichenbach 1853; Britton 1978]
  - Automolius angustulus (Burmeister, 1855) [by monotypy; MAMU] [=Automolus angustulus Burmeister, 1855]
- Bryantella Britton, 1957
  - Bryantella castanea Britton, 1957 [by monotypy; BMNH]
- Maechidinus Lea, 1919 [LR Britton 1957]
  - Maechidinus latericollis Lea, 1919 [by subsequent designation; Britton 1957; SAMA]
- Phyllochlaenia Blanchard, 1846 [Britton 1957; LR Britton 1957] [=Caulobius Le Guillou, 1844] [not Duponchel 1838] [=Deuterocaulobius Dalle Torre, 1912] [unnecessary n.n. pro Caulobius] [=Junkia Dalle Torre, 1913] [new name for Trichoderma; with Philochlaenia Moser 1913b] [=Philochlaenia Blanchard, 1845] [emend.] [=Trichoderma Nonfried, 1894] [not Stephens; Dalle Torre 1913]
  - Phyllochlaenia ceylanica (Nonfried, 1894) [by monotypy; ??] [=Trichoderma ceylanica Nonfried, 1894]
  - Phyllochlaenia rufescens Blanchard, 1846 [by monotypy; MNHP]
  - Phyllochlaenia villosus (Le Guillou, 1844) [by monotypy; MNHP] [=Caulobius villosus Le Guillou, 1844]

Tribù Chasmatopterini Lacordaire, 1856
- Chasmatopterus Latreille, 1825 [LR Baraud 1965; type sp. ?]
- Chnaunanthus Burmeister, 1844 [LR Saylor 1937] [=Acratus Horn, 1867] [not Lacordaire 1867; Dalle Torre 1912] [=Pseudacratus Dalle Torre, 1912] [Saylor 1937b]
  - Chnaunanthus discolor Burmeister, 1844 [by monotypy; MLUH]
  - Chnaunanthus flavipennis (Horn, 1867) [by monotypy; MCZC]

                                       [=Acratus flavipennis Horn, 1867]

Tribù Comophorini Britton, 1957 [=Comophini Britton, 1978]
- Comophorina Strand, 1928 [n. n. pro Comophorus]

                                  [=Comophorus Blanchard, 1851] [not Agassiz 1846; Strand 1928]
                                  [=Comophus Britton, 1978] [n.n. pro Comophorus in error Britton 1987]

- - Comophorina testaceipennis (Blanchard, 1851) [by monotypy; MNHN]

                                       [=Comophorus testaceipennis Blanchard, 1851]

Tribù Diphucephalini Britton, 1957
- Cunderdinia Lea, 1916
  - Cunderdinia variabilis Lea, 1916 [by monotypy; WAMP]
- Diphucephala Lepeletier & Serville, 1825 [LR Blackburn 1906]
  - Diphucephala sericea Kirby, 1818 [by monotypy; Silfverberg 1984; BMNH]

Tribù Heteronycini Britton, 1957 [key to genera Britton 1988]
- Acheilo Britton, 1988
  - Acheilo capitalis (Blackburn, 1910) [by original designation; BMNH]

                                       [=Heteronyx capitalis Blackburn, 1910]

- Callabonica Blackburn, 1895
  - Callabonica propria Blackburn, 1895 [by monotypy; BMNH]
- Cubidens Britton, 1992 [n.n. pro Paraheteronyx]

                                  [=Paraheteronyx Britton, 1988] [preoccupied]

- - Cubidens unguiculata (Burmeister, 1855) [by original designation; MLUH]

                                       [=Heteronyx unguiculata Burmeister, 1855]

- Eurychelus Blanchard, 1850
  - Eurychelus marmoratus Blanchard, 1850 [by monotypy; MNHN]
- Heteronyx Guerin-Meneville, 1831 [LR Blackburn 1911]

                                  [=Anacheirotus Blackburn, 1900]
                                  [=Cotidia Boisduval, 1835] [??]
                                  [=Hostilina Blanchard, 1851] [??]
                                  [=Silopa Erichson, 1842] [??]

- - Heteronyx australis Guerin-Meneville, 1831 [by monotypy; ?MNHN]
  - Heteronyx australis (Boisduval, 1835) [by monotypy; MNHN]

                                       [=Cotidia australis Boisduval, 1835]

- - Heteronyx glabrata (Erichson, 1842) [present designation; ZMHB]

                                       [=Silopa glabrata Erichson, 1842]

- - Heteronyx inornatus (Blackburn, 1900) [by monotypy; SAMA]

                                       [=Anacheirotus inornatus Blackburn, 1900]

- - Heteronyx praecox (Erichson, 1842) [by monotypy; ZMHB]

                                       [=Silopa praecox Erichson, 1842]

- Menigionyx Arrow, 1941a
  - Menigionyx laeviscutum Arrow, 1941a [by original designation; BMNH]
- Neoheteronyx Blackburn, 1890
  - Neoheteronyx lividus Blackburn, 1892 [by monotypy; SAMA]
- Nepytis Erichson, 1842
  - Nepytis russula Erichson, 1842 [by monotypy; ZMHB]
- Odontotonyx Macleay, 1871
  - Odontotonyx brunneipennis (Macleay, 1871) [by monotypy; MAMU]

                                       [=Odontolonyx brunneipennis Macleay, 1871]

- Paraheteronyx Moser, 1924a
  - Paraheteronyx quadriflabellatus Moser, 1924a [by monotypy; ZMHB]
- Proborhinus Britton, 1988
  - Proborhinus cornutus (Blackburn, 1910) [by monotypy; BMNH]

                                       [=Heteronyx cornutus Blackburn, 1910]

- Pseudoheteronyx Blackburn, 1892 [LR Britton 1988]
  - Pseudoheteronyx baldiensis (Blackburn, 1892) [by original designation; BMNH]

                                       [=Heteronyx baldiensis Blackburn, 1892]

- Webbella Britton, 1988
  - Webbella firmus (Blackburn, 1909) [by original designation; BMNH]

                                       [=Heteronyx firmus Blackburn, 1909]

Tribù Hopliini [?; where is Hoplia Illiger 1803 ?]
Tribù IncertaeSedis
- Anonetus Erichson, 1847 [Dalle Torre 1912; nomen nudum]
- Holophylla Erichson, 1847 [Britton 1978; nomen inquirendum]
- Hoplorida Moser, 1918 [nomen inquirendum]
  - Webbella congoana (Moser, 1918) [by monotypy; ZMHB; nomen inquirendum] [=Hoplorida congoana Moser, 1918]
- Tryssus Erichson, 1847 [Dalle Torre 1912; nomen nudum]
- Acoma Casey, 1889
  - Acoma brunnea Casey, 1889 [by monotypy; USNM]
- Alosimus [see Muche 1962]
- Conebius Fuavel, 1903
  - Conebius carinipennis Fauvel, 1903 [by monotypy; ??]
- Lacris Fairmaire & Germaine, 1860
  - Lacris dilutipes Fairmaire & Germaine, 1860 [by monotypy; ??]
- Metascelis Westwood, 1842 [see Allsopp 1989] TYPE SPECIES Metascelis flexilis Westwood, 1842 [by monotypyost]
- Podolasia Harold, 1869 [LR Howden 1954] [=Lasiopus Leconte, 1856] [not Schoenherr ??; Harold 1869]
  - Podolasia ferruginea (LeConte, 1856) [by monotypy; MCZC] [=Lasiopus ferruginea LeConte, 1856]

Tribù Liparetrini Burmeister, 1855 [=Allarini Britton, 1955b] [Britton 1980] [=Colpochilini Britton, 1957] [Britton 1980; key to the genera Britton 1990; ???]
- Allara Britton, 1955b
  - Allara insularis Britton, 1955b [by monotypy; WAMP]
- Allophyllus Fauvel, 1903
  - Allophyllus tetraphyllus Fauvel, 1903 [by monotypy; ??]
- Anacanthodes Britton, 1990 [n.n. pro Anacanthopus] [=Anacanthopus Blackburn, 1898] [preocc. ]
  - Anacanthodes inermis (Blackburn, 1898) [by monotypy; SAMA][=Anacanthopus inermis Blackburn, 1898]
- Aneucomides Blackburn, 1898
  - Aneucomides coloratus Blackburn, 1898 [by monotypy; BMNH]
- Aphanesia Britton, 1990
  - Aphanesia greyi Britton, 1990 [by monotypy; BMNH]
- Astibicola Britton, 1990
  - Astibicola bicolor Britton, 1990 [by monotypy; SAMA]
- Biphyllocera White, 1841
  - Biphyllocera kirbyana White, 1841 [by monotypy; BMNH]
- Cheilo Britton, 1990
  - Cheilo liparetroides Britton, 1990 [by monotypy; ANIC]
- Colobostoma Blanchard, 1850 [=Platydesmus Macleay, 1887] [not Lucas 1843; Britton 1990][=Pteroplatydesmus Dalle Torre, 1912] [n.n. pro Platydesmus; Britton 1990]
  - Colobostoma rufipennis (Boisduval, 1835) [by monotypy; MNHP] [=Sericesthis rufipennis Boisduval, 1835]
  - Colobostoma sulcipennis (Macleay, 1887) [by subsequent designation; Blackburn 1907; SAMA] [=Platydesmus sulcipennis Macleay, 1887]
- Colpochila Erichson, 1843 [LR Britton 1986] [=Haplonycha Blanchard, 1851] [Britton 1986]
  - Colpochila gouldii (Hope, 1842) [by monotypy; OXUM] [=Sericesthis gouldii Hope, 1842]
  - Colpochila obesa (Boisduval, 1835) [by subsequent designation; Blanchard 1853] [=Melolontha obesa Boisduval, 1835]
- Costelytra Given, 1952 [LR Given 1966]
  - Costelytra zealandica (White, 1844) [by original designation; BMNH] [=Rhisotrogus zealandica White, 1844]
- Costleya Broun, 1893
  - Costleya discoidea Broun, 1893 [by monotypy; ??]
- Dikellites Britton, 1990
  - Dikellites abditus Britton, 1990 [by monotypy; ANIC]
- Dysphanochila Blackburn, 1898
  - Dysphanochila pilosipennis Blackburn, 1898 [by monotypy; BMNH]
- Engyopsina Britton, 1990 [n.n. pro Engyops] [=Engyops Blackburn, 1898] [preoccupied]
  - Engyopsina spectans (Blackburn, 1898) [by monotypy; SAMA] [=Engyops spectans Blackburn, 1898]
- Frenchella Blackburn, 1892
  - Frenchella lubrica Blackburn, 1892 [by monotypy; SAMC]
- Glossocheilifer Blackburn, 1898
  - Glossocheilifer labialis Blackburn, 1898 [by monotypy; SAMC]
- Hadropechys Britton, 1990
  - Hadropechys victoriae (Blackburn, 1897) [by monotypy; BMNH][=Pachygastra victoriae Blackburn, 1897]
- Ictigaster Britton, 1986
  - Ictigaster ruficollis (Lea, 1917) [by monotypy; SAMA] [=Haplonycha ruficollis Lea, 1917]
- Leonotus Britton, 1986
  - Leonotus pilosicollis (Lea, 1930) [by monotypy; SAMA] [=Haplonycha pilosicollis Lea, 1930]
- Liparetrus Guerin-Meneville, 1831 [LR Britton 1980] [=Liparetra Hope, 1837]
  - Liparetrus discipennis Guerin-Meneville, 1831 [by monotypy; ?MNHP; see Sherborne & Woodward 1906 on the publication date 1831 = 1838]
  - Liparetrus sylvicola (Fabricius, 1775) [by original designation; BMNH] [=Melolontha sylvicola Fabricius, 1775]
- Macleayella Britton, 1990 [n.n. pro Macleayia] [=Macleayia Blackburn, 1888] [preoccupied; Blackburn 1905]
  - Macleayella singularis (Blackburn, 1888) [by subsequent designation; Britton 1990; BMNH] [=Macleayia singularis Blackburn, 1888]
- Neso Blackburn, 1898
  - Neso usta Blackburn, 1898 [by subsequent designation; Britton 1990; SAMA]
- Nosphisthis Blackburn, 1898
  - Nosphisthis parvicornis Blackburn, 1898 [by monotypy; SAMA]
- Odontria White, 1846 [LR Given 1952]
  - Odontria striata White, 1846 [by ??; BMNH]
- Pachygastra Germar, 1848
  - Pachygastra tasmanica Germar, 1848 [by monotypy; MLUH]
- Parasciton Britton, 1990
  - Parasciton inermis Britton, 1990 [by monotypy; ANIC]
- Paronyx Britton, 1990
  - Paronyx setifera Britton, 1990 [by monotypy; ANIC]
- Petinopus Blackburn, 1898
  - Petinopus aegrotus Blackburn, 1898 [by monotypy; BMNH]
- Photyna Brenske, 1897b
  - Photyna rugicollis Brenske, 1897b [by present designation; ??]
- Poecilodiscus Broun, 1895 [LR Given 1952]
  - Poecilodiscus pulcher Broun, 1895 [by monotypy; BMNH]
- Prodontria Broun, 1904 [LR Given 1960] [=Lewisiella Broun, 1909] [Given 1952]
  - Prodontria lewissi Broun, 1904 [by montypy; BMNH]
  - Prodontria modesta (Broun, 1909) [by subsequent designation; Given 1952; BMNH] [=Lewisiella modesta Broun, 1909]
- Scitaloides Britton, 1990
  - Scitaloides malanda Britton, 1990 [by monotypy; ANIC]
- Sciton Blackburn, 1892
  - Sciton ruber (Blackburn, 1892) [by monotypy; SAMA][=Scition ruber Blackburn, 1892]
- Scythrodes Broun, 1886 [LR Given 1952]
  - Scythrodes squalidus Broun, 1881
- Sericospilus Sharp, 1882 [LR Given 1952] [=Eusoma White, 1846] [preoccupied; Sharp 1882]
  - Sericospilus rossii (White, 1846) [by original designation; ??] [=Eusoma rossii White, 1846]
- Stenochelyne Britton, 1990
  - Stenochelyne noctis Britton, 1990 [by monotypy; ANIC]
- Teluroides Britton, 1990
  - Teluroides suturalis (Lea, 1920) [by original designation; SAMA] [=Telura suturalis Lea, 1920]
- Xyridea Britton, 1990
  - Xyridea hirticollis (Blackburn, 1912) [by monotypy; BMNH] [=Aneucomides hirticollis Blackburn, 1912]
Tribù Macrodactylini Burmeister, 1855 [subfamily?] [=Dichelonyciini Ritcher, 1966] [???]
- Acanthosternum Philippi, 1861
  - Acanthosternum splendens Philippi, 1861 [by monotypy; ??]
- Agaocnemis Moser, 1918
  - Agaocnemis pruina Moser, 1918 [by monotypy; ZMHB]
- Alvarinus Blanchard, 1851
  - Alvarinus hilarii Blanchard, 1851 [present designation; MNHN]
- Ancistroma Curtis, 1835 [LR Arrow 1913]
  - Ancistroma klugi Curtis, 1835 [by monotypy; BMNH]
- Anomalochilus Blanchard, 1850
  - Anomalochilus singularis Blanchard, 185 [by monotypy; MNHN]
- Anomonyx Saylor, 1940a [=Anomalonyx Moser, 1922] [not Weise 1903; Saylor 1940]
  - Anomonyx uruguayensis (Moser, 1922) [by monotypy; ZMHB] [=Anomalonyx uruguayensis Moser, 1922]
- Anoplosiagum Blanchard, 1850
  - Anoplosiagum sulcatum Blanchard, 1850 [by present designation; MNHN]
- Apterodemidea Gutierrez Alonso, 1952
  - Apterodemidea paraguayensis (Arrow, 1903) [by monotypy; BMNH] [=Apterodema paraguayensis Arrow, 1903]
- Astaenoplia Martinez, 1957
  - Astaenoplia miserabilis Martinez, 1957 [by monotypy; AMIC]
- Astaenosiagum Martinez, 1957
  - Astaenosiagum longipes (Philippi, 1861) [by monotypy; MNNC] [=Schizochelus longipes Philippi, 1861]
- Aulanota Moser, 1924a
  - Aulanota sulcipennis Moser, 1924a [by monotypy; ZMHB]
- Barybas Burmeister, 1855 [=Microcrania Burmeister, 1855] [fide Dalla Torre 1912]
  - Barybas compacta (Erichson, 1847) [present designation; MLUH] [=Philochlaenia compacta Erichson, 1847]
  - Barybas volvulus Burmeister, 1855 [by monotypy; MLUH]
- Byrasba Harold, 1869
  - Byrasba volvulus (Burmeister, 1855) [by monotypy; MLUH] [=Barybas volvulus Burmeister, 1855]
- Calodactylus Blanchard, 1851 [=Doplia Burmeister, 1855] [fide Dalla Torre 1912]
  - Calodactylus sulphurea (Burmeister, 1855) [present designation; MLUH] [=Doplia sulphurea Burmeister, 1855]
  - Calodactylus tibialis (Blanchard, 1851) [by monotypy; MNHN] [=Calodactylys tibialis Blanchard, 1851]
- Canestera Saylor, 1938a
  - Canestera marshalli Saylor, 1938a [by monotypy; ??]
- Ceraspis Lepeletier & Serville, 1825 [LR Frey 1962]

SUBGENUS Ceraspis Lepeletier & Serville, 1825

- - Ceraspis pruinosa Lepeletier & Serville, 1825 [by subsequent designation; ??; ??]

SUBGENUS Isoceraspis Kolbe, 1911 [=Faula Blanchard, 1950] [Burmeister 1855]

- - Ceraspis cornuta (Blanchard, 1850) [present designation; MNHN] [=Faula cornuta Blanchard, 1850]
  - Ceraspis duckei Kolbe, 1911 [by ??; ??]
- Ceratolontha Arrow, 1948a
  - Ceratolontha venezuelae Arrow, 1948a [by monotypy; BMNH]
- Chariodactylus Moser, 1919
  - Chariodactylus chacoensis Moser, 1919 [present designation; ZMHB]
- Chariodema Blanchard, 1850
  - Chariodema virescens (Blanchard, 1850) [present designation; MNHN] [=Philochlaenia virescens Blanchard, 1850]
- Clavipalpus Laporte, 1832 [=Amphicrania Burmeister, 1855] [fide Dalla Torre 1912]
  - Clavipalpus dejeani Laporte, 1832 [by monotypy; ??]
  - Clavipalpus ursina (Burmeister, 1855) [present designation; MLUH] [=Amphicrania ursina Burmeister, 1855]
- Clitocephala Burmeister, 1855
  - Clitocephala pellucens Burmeister, 1855 [by monotypy; MLUH]
- Coenonycha Horn, 1876 [LR Cazier & McClay 1943; Evans & Smith 1986]
  - Coenonycha rotundata (LeConte, 1856) [by subsequent designation; ??; MCZC] [=Dichelonyx rotundata LeConte, 1856]
- Corminus Burmeister, 1855
  - Corminus canescens Burmeister, 1855 [present designation; MLUH]
- Cremastodus Solier, 1851
  - Cremastodus pubescens Solier [present designation]
- Ctenotis Burmeister, 1855
  - Ctenotis obesa Burmeister, 1855 [by monotypy; MLUH]
- Dasyus Lepeletier & Serville, 1825
  - Dasyus collaris Lepeletier & Serville, 1825 [by monotypy; ??]
- Demodema Blanchard, 1850
  - Demodema fallax Blanchard, 1850 [present designation; MNHN]
- Diaphylla Erichson, 1847
  - Diaphylla hispida Erichson, 1847 [by monotypy; ZMHB]
- Dichelonyx Harris, 1827 [LR Saylor 1945] [=Dichelonycha Kirby, 1837] [Pate 1946; type sp.?]
  - Dichelonyx linearis (Gyllenhal, 1817) [junior synonym of M. elongata Fabricius 1792; by original designation; ??][=Melolontha linearis Gyllenhal, 1817]
- Dicrania Lepeletier & Serville, 1825 [LR Frey 1972]
  - Dicrania nigra Serville, 1825 [by monotypy; ??]
- Diphycerus Deyrolle & Fairmaire, 1878
  - Diphycerus davidis Deyrolle & Fairmaire, 1878 [by monotypy; MNHN]
- Diphydactylus Thomson, 1858
  - Diphydactylus singularis, 1858 [by monotypy; ??]
- Eubarybas Gutierrez Alonso, 1952
  - Eubarybas asper Gutierrez Alonso, 1952 [by monotypy; ??]
- Gama Blanchard, 1850[=Harpodactyla Burmeister, 1855]
  - Gama grandicornis (Burmeister, 1855) [present designation; MLUH] [=Harpodactyla grandicornis Burmeister, 1855]
  - Gama relucens Blanchard, 1850 [present designation; MNHN]
- Gastrohoplus Moser, 1921b
  - Gastrohoplus mirabilis Moser, 1921b [by monotypy; ZMHB]
- Gymnopyge Linell, 1895
  - Gymnopyge hopliaeformis Linell, 1895 [by original designation; USNM]
- Hadrocerus Guerin-Meneville, 1830
  - Hadrocerus castaneipennis Guerin-Meneville, 1830 [by monotypy; ??]
- Hamatoplectris Frey, 1967
  - Hamatoplectris rosettae Frey, 1967 [by original designation; ZSMC]
- Hercites Burmeister, 1855
  - Hercites pygmaea Burmeister, 1855 [by monotypy; MLUH]
- Hieritis Burmeister, 1855
  - Hieritis macrocera Burmeister, 1855 [by monotypy; MLUH]
- Homalochilus Blanchard, 1851
  - Homalochilus niger Blanchard, 1851 [present designation; MNHN]
- Homioliogenys Gutierrez Alonso, 1952
  - Homioliogenys tarsalis (Moser) [by monotypy; ZMHB; ??] [=Liogenys tarsalis Moser]
- Hyperius Deyrolle & Fairmaire, 1878
  - Hyperius davidis Deyrolle & Fairmaire, 1878 [present designation; MNHN]
- Isonychus Mannerheim, 1829 [LR South American spp. Frey 1970]
  - Isonychus sulphureus Mannerheim, 1829 [present designation; ??]
- Issacaris Fairmaire, 1889 [LR Gutierrez Alonso 1952]
  - Issacaris petalophora Fairmaire, 1889 [by monotypy; MNHN]
- Liogenys Guerin-Meneville, 1830
  - Liogenys castaneus Guerin -Meneville, 1830 [senior ?homonym of Liogenys castaneus Curtis 1845; junior synonym of Melolontha palpalis castanea Escscholtz 1822; by monotypy; ??]
- Macrodactylus Latreille, 1825 [LR of Mexican spp. Carrillo & Gibson 1960; type sp.?]
- Mallotarsus Blanchard, 1850
  - Mallotarsus spadiceus Blanchard, 1850 [by monotypy; MNHN]
- Manodactylus Moser, 1919a
  - Manodactylus gaujoni Moser, 1919a [by monotypy; ZMHB]
- Manonychus Moser, 1919b
  - Manonychus unguicularis Moser, 1919b [by monotypy; ZMHB]
- Manopus Laporte, 1840
  - Manopus biguttatus Laporte, 1840 [by monotypy; ??]
- Metaceraspis Frey, 1962
  - Metaceraspis fukiensis Frey, 1962 [by monotypy; ZSMC]
- Oedichira Burmeister, 1855
  - Oedichira pachydactyla Burmeister, 1855 [present designation; MLUH]
- Pachylotoma Blanchard, 1850
  - Pachylotoma viridis Blanchard, 1850 [by monotypy; MNHN]
- Pacuvia Curtis, 1845
  - Pacuvia castanea Curtis, 1845 [by monotypy; ??]
- Pectinosoma Arrow, 1913
  - Pectinosoma elongata Arrow, 1913 [by monotypy; BMNH]
- Plectris Lepeletier & Serville, 1825

SUBGENUS Philochlaenia Blanchard, 1851 [see Deuterocaulobius; LR Frey 1967; type sp.?] SUBGENUS Plectris Lepeletier & Serville, 1825 [=Euryaspis Blanchard, 1851] [fide Dalla Torre 1912] [=Pseudoserica Guerin-Menenville, 1830] [Dalle Torre 1912; not syn. Arrow 1948a; type sp.?]

- - Plectris gaudaudii (Blanchard, 1851) [by monotypy; MNHN] [=Euryaspis gaudaudii Blanchard, 1851]
  - Plectris tomentosa Lepeletier & Serville, 1825 [by monotypy; ??]
- Pristerophora Harold, 1869 [=Prionophora Solier, 1851] [not Westwood 1848; Harold 1869]
  - Pristerophora picipennis (Solier, 1851) [by monotypy; ??] [=Prionophora picipennis Solier, 1851]
- Pseudodicrania Gutierrez Alonso, 1950a
  - Pseudodicrania aeneobrunnea (Philippi, 1861) [by monotypy; MNNC][=Dicrania aeneobrunnea Philippi, 1861]
- Pseudohercitis Moser, 1921b
  - Pseudohercitis viridiaenea Moser, 1921b [by original designation; ??]
- Pseudoisonychus Frey, 1971a
  - Pseudoisonychus vietnamensis Frey, 1971a
- Rhinaspis Perty, 1833
  - Rhinaspis schranki Perty, 1833 [junior synonym of Melolontha aenea Billberg 1820; fide Burmeister 1855; by monotypy; ??]
- Rhinaspoides Moser, 1919b
  - Rhinaspoides aeneofusca (Moser, 1919b) [by monotypy; ZMHB] [=Rhinapoides aeneofusca Moser, 1919b]
- Schizochelus Blanchard, 1850
  - Schizochelus bicoloripes Blanchard, 1850 [by monotypy; MNHN]
- Ulomenes Blanchard, 1851
  - Ulomenes hypocrita Mannerheim, 1829 [by monotypy; MNHN]
- Xenoceraspis Arrow, 1920
  - Xenoceraspis dispar Arrow, 1920 [by monotypy; BMNH]
- Zabacana Saylor, 1946
  - Zabacana brevinigrans Saylor, 1946 [by monotypy; CASC]
Tribù Maechidini Burmeister, 1855 [key to the genera Britton 1957]
- Epholcis Waterhouse, 1875a [LR Britton 1957]
  - Epholcis divergens Waterhouse, 1875a [by monotypy; BMNH]
- Harpechys Britton, 1957
  - Harpechys chilo Britton, 1957 [by monotypy; SAMA]
- Maechidius Macleay, 1819 [LR Britton 1957]
  - Maechidius spurius (Kirby, 1818) [by monotypy; ?BMNH] [=Trox spurius Kirby, 1818]
- Microcoenus Britton, 1957
  - Microcoenus nanus Britton, 1957 [by monotypy; ANIC]
- Microthopus Burmeister, 1855
  - Microthopus castanopterus Burmeister, 1855 [by monotypy; MLUH]
- Paramaechidius Frey, 1969b
  - Paramaechidius pauxillus (Heller) [by original designation; ??] [=Maechidius pauxillus Heller]
- Termitophilus Britton, 1957
  - Termitophilus spadix Britton, 1957 [by monotypy; WAMP]
Tribù Melolonthini Burmeister, 1855 [North America-Hardy 1974; Australia-Britton 1978; MacLeay 1819??]
- Achranoxia Kraatz, 1888
  - Achranoxia koenigi (Brenske, 1888) [by monotypy; ??] [=Polyphylla koenigi Brenske, 1888]
- Adoretops Kraatz, 1883
  - Adoretops plexus (Zoubkoff, 1833) [by monotypy; ??] [=Melolontha plexus Zoubkoff, 1833]
- Aethiartrogus Decelle, 1982
  - Aethiartrogus clypeatus (Blanchard, 1851) [by original designation; MNHN] [=Rhizotrogus clypeatus Blanchard, 1851]
- Afrolepis Decelle, 1968 [LR Decelle 1968] [=Oligolepis Brenske, 1903a] [not Bleeker 1874; Decelle 1968; nomen nudum;= Stethaspis Arrow 1943]
  - Afrolepis pygidilalis (Brenske, 1903a) [by subsequent designation; Lepesme & Paulian 1944; ZMHB] [=Oligolepis pygidilalis Brenske, 1903a]
- Allothnonius Britton, 1978
  - Allothnonius brooksi Britton, 1978 [by original designation; ANIC]
- Amblonoxia Reitter, 1902
  - Amblonoxia quedenfeldti (Kraatz, 1894) [by monotypy; ZMHB] [=Oligophylla quedenfeldti Kraatz, 1894]
- Amphimallina Reitter, 1905
  - Amphimallina jenrichi Reitter, 1905 [by monotypy; ??]
- Amphimallon Le Peletier & Serville, 1825

SUBGENUS Amadotrogus Reitter, 1902

- - Amphimallon scutellaris (Lucas, 1849) [by monotypy; MNHN] [=Rhizotrogus scutellaris Lucas, 1849]

SUBGENUS Amphimallon Medvedev, 1951 [=Microdonta Hope, 1837] [Lacordaire 1856]

- - Amphimallon pini (Fabricius) [by original designation; OXUM?] [=Melolontha pini Fabricius]
  - Amphimallon solstitialis (Linnaeus, 1758) [by subsequent designation; Medvedev 1951; BMNH] [=Scarabaeus solstitialis Linnaeus, 1758]

SUBGENUS Dasytrogus Reitter, 1902 [as subgenus of Madatrogus Nikolaev 1976]

- - Amphimallon transcaspicus Brenske, 1886 [by monotypy; ??]

SUBGENUS Eriotrogus Reitter, 1902

- - Amphimallon erivanicus Reitter, 1902 [by monotypy; ??]

SUBGENUS Madotrogus Reitter, 1902 [*cited as genus Nikolaev 1976; ???]

- - Amphimallon glabricollis Reitter, 1888 [by original designation; ??]

SUBGENUS Micramphimallon Medvedev, 1951

- - Amphimallon mesopotamicus (Medvedev, 1951) [by monotypy; ??] [=Micramphimallon mesopotamicus Medvedev, 1951]
- Anartioschiza Kolbe, 1894 [LR Decelle 1975]
  - Anartioschiza cameruna Kolbe, 1894 [by subsequent designation; Decelle 1975; ZMHB]
- Anisopholis Moser, 1913b
  - Anisopholis squamulifera (Moser, 1913b) [by original designation; ZMHB] [=Schoenherria squamulifera Moser, 1913b]
- Anomalochela Moser, 1913a [= Lachnosterna Arrow 1948b; status reviewed by Khan& Ghai 1983]
  - Anomalochela bicolor (Brenske, 1892) [by original designation; ZMHB] [=Brahmina bicolor Brenske, 1892]
- Anoxia Laporte, 1832 [LR Dewailly 1944; key to subgenera Sabatinelli 1976; type sp.?]

SUBGENUS Anoxia Medvedev, 1951 SUBGENUS Archianoxia Miksic, 1959

- - Anoxia derelicta Desbrochers, 1873 [by original designation; ??]

SUBGENUS Mesanoxia Medvedev, 1951

- - Anoxia australis (Gyllenhal, 1817) [by original designation; ??] [=Melolontha australis Gyllenhal, 1817]

SUBGENUS Protanoxia Medvedev, 1951

- - Anoxia orientalis (Krynizky, 1832) [by original designation; ??] [=Melolontha orientalis Krynizky, 1832]
- Anoxoides Petrovitz, 1971
  - Anoxoides bytinskisalzi Petrovits, 1971 [by monotypy; ??]
- Antitrogus Burmeister, 1855 [LR Britton 1978; 1979]
  - Antitrogus tasmanicus Burmeister, 1855 [by monotypy; MLUH]
- Aplidia Hope, 1837 [LR Baraud 1988/89?] [=Haplidia Gemminger & Harold, 1869] [unjustified emendation]
  - Aplidia transversa (Fabricius, 1801) [by monotypy; BMNH] [=Melolontha transversa Fabricius, 1801]
- Apogonia Kirby, 1818

SUBGENUS Apogonia Kirby, 1818 [=Catagonia Kolbe, 1899] [Moser 1917a] [=Dichecephala Brenske, 1895] [Arrow 1940; =Catagonia Moser 1917a; note, Moser 1917 considered Catagonia a subgenus of Dichecephala]

- - Apogonia abysinnica (Brenske, 1895) [=Dichecephala abysinnica Brenske, 1895]
  - Apogonia gemellata Kirby, 1818 [by ??; a junior synonym of Melolontha rauca Fabricius 1781 now in Apogonia]
  - Apogonia nasalis Karsch, 1882 [by monotypy; ZMHB]

SUBGENUS Ceratogonia Kolbe, 1899 [type sp.?] SUBGENUS Metagonia Kolbe, 1899

- - Apogonia platypyge Kolbe, 1899 [present designation; ZMHB]

SUBGENUS Rhyncogonia Arrow, 1902

- - Apogonia acuminata Arrow, 1902 [by monotypy; BMNH; note, Arrow does not state "n. subg."]
- Aposchiza Brenske, 1903c
  - Aposchiza schultzi Brenske, 1903c [by monotypy; ZMHB]
- Articephala Moser, 1920
  - Articephala planifrons Moser, 1920 [by monotypy; ZMHB]
- Asactopholis Brenske, 1894b
  - Asactopholis gracilipes (Sharp, 1876) [by original designation; ??] [=Lepidiota gracilipes Sharp, 1876]
- Asaphomorpha Brancsik, 1893 [LR Dewailly 1950]
  - Asaphomorpha nigra (Nonfried, 1891) [by monotypy; MNHN] [=Encya nigra Nonfried, 1891]
- Asophrops Petrovitz, 1961
  - Asophrops jettmari Petrovitz, 1961 [by monotypy; ??]
- Asthenopholis Brenske, 1898 [LR Arrow 1917]
  - Asthenopholis transvaalensis (Brenske, 1898) [present designation; ZMHB] [=Astenopholis transvaalensis Brenske, 1898]
- Aulacoschiza Decelle, 1973b
  - Aulacoschiza triphylla Decelle, 1973b [by original designation; MRAC]
- Beriqua Peringuey, 1904
  - Beriqua modesta Peringuey, 1904 [by monotypy; SAMC]
- Brachylepis Kolbe, 1894
  - Brachylepis elephas (Gerstaecker, 1867) [by monotypy; ZMHU] [=Coniopholis elephas Gerstaecker, 1867]
- Brachyllus Brenske, 1896
  - Brachyllus ulcerosus Brenske, 1896 [present designation; ZMHU]
- Brachypholis Brenske, 1898b [=Epipholis Moser, 1917b] [Arrow 1943]
  - Brachypholis assiniensis (Moser, 1917b) [by monotypy; ZMHU] [=Epipholis assiniensis Moser, 1917b]
  - Brachypholis rothschildi Brenske, 1898b [by monotypy; ZMHU]
- Brahmina Blanchard, 1851

SUBGENUS Anoxiella Reitter, 1902 [=Schistocamenta Seidlitz, 1905] [err. pro Schistocometa]

- - Brahmina brenskei Reitter, 1900 [by monotypy; ZMHB]
  - Brahmina senescens Frivaldszky, 1889 [by subsequent designation; Medvedev 1951; ??]

SUBGENUS Brahmina Blanchard, 1851

- - Brahmina comata Blanchard, 1851 [by montypy; MNHN]

SUBGENUS Brahminella Medvedev, 1951

- - Brahmina agenlla Faldermann, 1835 [by original designation; ??]

SUBGENUS Brahminula Medvedev, 1951 [=Schistocometa Brenske, 1903] [Reitter 1904]

- - Brahmina alatavica Semenov-Tian-Shanskii, 1925 [by monotypy; ??]
- Butozania Miksic, 1955 [subgenus of Rhizotrogus; Kantardschieva-Minkova 1958]
  - Butozania macedonicus (Miksic, 1955) [by monotypy; ??] [=Rhizotrogus macedonicus Miksic, 1955]
- Byrsalepis Brenske, 1898
  - Byrsalepis mikindana Brenske, 1898 [present designation; ZMHB]
- Cephaloschiza Moser, 1920 [LR Decelle 1979]
  - Cephaloschiza clypealis Moser, 1920 [by monotypy; ZMHB]
- Chaetocosmetes Moser, 1917b
  - Chaetocosmetes javanus Moser, 1917 [present designation; ZMHB]
- Chariochilus Brenske, 1898b
  - Chariochilus metallescens Brenske, 1898b [by monotypy; ZMHB]
- Cheronia Croissandaeu, 1896
  - Cheronia bonnemaisoni Croissandaeu, 1896 [by monotypy; ??]
- Chiloschiza Moser, 1924a
  - Chiloschiza clypealis Moser, 1924a [by monotypy; ZMHB]
- Chilotrogus Reitter, 1905
  - Chilotrogus panotrogoides Reitter, 1905 [by monotypy; ??]
- Chioneosoma Kraatz, 1881 [LR Medvedev 1966]

SUBGENUS Aleucolomus Reitter, 1902

- - Chioneosoma subporosa Reitter, 1902 [by monotypy; ??]

SUBGENUS Aleucoschema Semenov-Tian-Shanskii, 1935

- - Chioneosoma vulpina (Gyllenhal, 1817) [by subsequent designation; Medvedev 1951; ??] [=Melolontha vulpina Gyllenhal, 1817]

SUBGENUS Chioneosoma Kraatz, 1881 [Reitter 1902]

- - Chioneosoma pulvereum (Knoch, 1801) [by subsequent designation; Medvedev 1951; ??] [=Melolontha pulvereum Knoch, 1801]

SUBGENUS Chionotrogus Reitter, 1902

- - Chioneosoma pulverea (Knoch, 1801) [by subsequent designation; Medevedv 1951; ??] [=Melolontha pulverea Knoch, 1801]

SUBGENUS Leucolasium Semenov-Tian-Shanskii, 1935

- - Chioneosoma gorilla (Brenske, 1886) [by subsequent designation; Medvedev 1951; ZMHB] [=Rhizotrogus gorilla Brenske, 1886]

SUBGENUS Leucoschema Semenov-Tian-Shanskii, 1935

- - Chioneosoma baadeni (Brenske, 1886) [by subsequent designation; Medvedev 1951; ZMHB] [=Rhizotrogus baadeni Brenske, 1886]

SUBGENUS Leucoschesis Semenov-Tian-Shanskii, 1935

- - Chioneosoma turcomanum Brenske, 1888 [by subsequent designation; Medvedev 1951; ZMHB]
- Clypeasta Fairmaire, 1904a [LR Dewailly 1950] [=Clypearia Fairmaire, 1903] [preoccupied; Fairmaire 1904a]
  - Clypeasta pruinosa (Fairmaire, 1903) [by monotypy; MNHN] [=Clypearia pruinosa Fairmaire, 1903]
- Cnemoschiza Decelle, 1979
  - Cnemoschiza tibialis Decelle, 1979 [by monotypy; MZUF]
- Cochliotis Kolbe, 1894
  - Cochliotis melolonthoides (Kolbe, 1894) [by monotypy; ZMHB]

                                       [=Coniopholis melolonthoides Kolbe, 1894]

- Cochliotodes Burgeon, 1946b
  - Cochliotodes babaulti Burgeon, 1946b [by monotypy; ??]
- Coelogenia Moser, 1913b
  - Coelogenia flavida Moser, 1913b [by monotypy; ZMHB]
- Coniopholis Erichson, 1847
  - Coniopholis lepidiota Burmeister, 1855 [by subsequent monotypy; MLUH]
- Coniotrogus Brenske, 1902b
  - Coniotrogus abdominalis Brenske, 1902b [by monotypy; ZMHB]
- Crepischiza Brenske, 1903c
  - Crepischiza usambarae Bresnke, 1903 [by monotypy; ZMHB]
- Cryphaeobius Kraatz, 1882
  - Cryphaeobius brunneus Kraatz, 1882 [by monotypy; ZMHB]
- Cryptotrogus Kraatz, 1888
  - Cryptotrogus weisei Kraatz, 1888 [by monotypy; ZMHB]
- Cyphochilus Waterhouse, 1867
  - Cyphochilus candida (Olivier, 1789) [by subsequent designation; Medvedev 1951; ??]

                                       [=Melolontha candida Olivier, 1789]

- Cyphonotus Fischer, 1824 [LR Medvedev 1951]
  - Cyphonotus testaceus (Pallas, 1781) [by subsequent designation; Medevedev 1951; ??]

                                       [=Scarabaeus testaceus Pallas, 1781]

- Cyphonoxia Reitter, 1889 [LR Medvedev 1951]
  - Cyphonoxia praestabilis Reitter, 1889 [by monotypy; ??]
- Dasylepida Moser, 1913b
  - Dasylepida fissa Moser, 1913b [by present designation; ZMHB]
- Dermolepida Arrow, 1941 [n.n.pro Lepidoderma; LR Britton1978]

                                  [=Lepidoderma Waterhouse, 1875b] [not Reuss 1855]

- - Dermolepida albohirtum (Waterhouse, 1875b) [by monotypy; BMNH]

                                       [=Lepidoderma albohirtum Waterhouse, 1875b]

- Diacucephalus Dewailly, 1950
  - Diacucephalus vadoni Dewailly, 1950 [by original designation; MNHN]
- Dicheloscema Moser, 1924
  - Dicheloscema corporaali (Moser, 1924) [by monotypy; ZMHB]

                                       [=Dicheloschema corporaali Moser, 1924]

- Dinacoma Casey, 1889 [LR Hardy 1974]
  - Dinacoma marginata Casey, 1889 [by monotypy; USNM]
- Diplotaxis Kirby, 1837 [LR Vaurie 1958;1960]

                                  [=Alobus LeConte, 1856] [Vaurie 1958]
                                  [=Diazus Leconte, 1860] [Vaurie 1958]
                                  [=Orsonyx LeConte, 1856] [Fall 1909]

- - Diplotaxis anxius (LeConte, 1856) [by monotypy; MCZC]

                                       [=Orsonyx anxius LeConte, 1856]

- - Diplotaxis fulvus (LeConte, 1856) [by monotypy; MCZC]

                                       [=Alobus fulvus LeConte, 1856]

- - Diplotaxis rudisLeConte, 1860) [by monotypy; MCZC]

                                       [=Diazus rudisLeConte, 1860]

- - Diplotaxis tristis Kirby, 1837 [by monotypy; BMNH]
- Empecta Erichson, 1847 [LR Dewailly 1950]
  - Empecta obsoleta Blanchard, 1851 [by subsequent designation; Dewailly 1950; MNHN]
- Empectida Moser, 1914b
  - Empectida vethi Moser, 1914 [by monotypy; ZMHB]
- Empectoides Dewailly, 1950
  - Empectoides gentilis (Blanchard, 1851) [by original designation; MNHN]

                                       [=Empecta gentilis Blanchard, 1851]

- Empycastes Gerstaecker, 1871
  - Empycastes coronatus Gerstaecker, 1871 [by monotypy; ZMHB]
- Enaria Erichson, 1847
  - Enaria melanictera (Klug, 1833) [by subsequent monotypy; Blanchard 1851; ZMHB]

                                       [=Melolontha melanictera Klug, 1833]

- Encya Blanchard, 1845 [LR Dewailly 1950; note, Burm. 1855 cites Dejean 1833]
  - Encya apicalis Blanchard, 1851 [by subsequent designation; Dewailly 1950; MNHN]
- Energetia Dalle Torre, 1912

                                  [=Phila Brenske, 1897b] [not Motschulsky 1864; Dalle Torre 1912]

- - Energetia amboinae (Brenske, 1897b) [by monotypy; ZMHB]

                                       [=Phila amboinae Brenske, 1897b]

- Enthora Erichson, 1847 [LR Dewailly 1950]
  - Enthora chlorodera Blanchard, 1851 [by subsequent monotypy; MNHN]
- Entyopsis Kolbe, 1894
  - Entyopsis impressa Kolbe, 1894 [by original desgnation; ZMHB]
- Entypophana Moser, 1913
  - Entypophana biapicata Moser, 1913 [present designation; ZMHB]
- Eremotrogus Kolbe, 1914
  - Eremotrogus pruinosus Kolbe, 1914 [by monotypy; ZMHB]
- Eulepidopsis Burgeon, 1946a
  - Eulepidopsis wilsoni Burgeon, 1946a [by monotypy; ??]
- Eupegylis Duvivier, 1892
  - Eupegylis confusa Duvivier, 1892 [by monotypy; ??]
- Euranoxia Semenov-Tian-Shanskii, 1890
  - Euranoxia valida (Semenov, 1890) [by monotypy; ??]

                                       [=Euronoxia valida Semenov, 1890]

- Eutrichesis Waterhouse, 1882 [LR Dewailly 1950]
  - Eutrichesis punctatus Waterhouse, 1882 [present designation; BMNH]
- Exolontha Reitter, 1902
  - Exolontha umbraculata (Burmeister, 1855) [by monotypy; MLUH]

                                       [=Melolontha umbraculata Burmeister, 1855]

- Exopholis Motschulsky, 1859
  - Exopholis birmanica Motschulsky, 1959 [by monotypy; ??]
- Genyoschiza Moser, 1917b

                                  [=Atys Reiche, 1850] [not de Montfort 1810; =Genyoschiza Arrow 1944; preocc.Strand 1942]
                                  [=Atysilla Strand, 1942]

- - Genyoschiza samenensis (Reiche, 1850) [by monotypy; BMNH]

                                       [=Atys samenensis Reiche, 1850]

- - Genyoschiza tarsalis Moser, 1917b [by monotypy; ZMHB]
- Geotrogus Guerin-Meneville, 1842
  - Geotrogus magagnosci Guerin-Meneville, 1842 [by monotypy; ??]
- Globulischiza Frey, 1974b
  - Globulischiza squamosa Frey, 1974b [by monotypy; ZSMC]
- Glyptoglossa Brenske, 1895
  - Glyptoglossa lurida (Brumeister, 1855) [by monotypy; MLUH]

                                       [=Schizonycha lurida Brumeister, 1855]

- Gnaphalostetha Reiche, 1856
  - Gnaphalostetha bonvoisini (Reiche, 1856) [by monotypy; ??]

                                       [=Gnaphalostoma bonvoisini Reiche, 1856]

- Grananoxia Brenske, 1890
  - Grananoxia annamensis (Fleutiaux, 1887) [by monotypy; ??]

                                       [=Granida annamensis Fleutiaux, 1887]

- Gymnogaster Blanchard, 1851 [preocc. Gronorius 1763; Brunnich 1788]
  - Gymnogaster buphthalmus Blanchard, 1851 [by monotypy; MNHN]
- Gymnoschiza Moser, 1914a [LR Decelle 1973a]
  - Gymnoschiza gracilipes (Brenske, 1898) [by original designation; ZMHB]

                                       [=Anartioschiza gracilipes Brenske, 1898]

- Hecistopsilus Kolbe, 1894
  - Hecistopsilus molitor (Kolbe, 1894) [present designation; ??]

                                       [=Hecistospilus molitor Kolbe, 1894]

- Heptelia Brenske, 1898b [LR Arrow 1944b]
  - Heptelia stripidea Brenske, 1898b [by monotypy; ZMHB]
- Heptophyllla Motschulsky, 1857
  - Heptophyllla picea (Motschulsky, 1857) [by monotypy; ??]

                                       [=Heptophylla picea Motschulsky, 1857]

- Heteroschiza Moser, 1917b
  - Heteroschiza plumata Moser, 1917b [by monotypy; ZMHB]
- Hexataenius Fairmaire, 1891
  - Hexataenius protensus Fairmaire, 1891 [by monotypy; MNHN]
- Hilyotrogus Fairmaire, 1886

                                  [=Melichrus Brenske, 1892a] [Arrow 1944; Frey 1975b]

- - Hilyotrogus kolbei (Brenske, 1892a) [by monotypy; ZMHB]

                                       [=Melichrus kolbei Brenske, 1892a]

- - Hilyotrogus unguicularis Fairmaire, 1886 [present designation; MNHN]
- Holisonycha Peringuey, 1904
  - Holisonycha mellila Peringeuy, 1904 [by monotypy; SAMC]
- Holochelus Reitter, 1889

SUBGENUS Holochelus Reitter, 1889 [Semenov 1905]

- - Holochelus subseriatus Reitter, 1889 [by monotypy; ??]

SUBGENUS Odontochelus Semenov, 1905

- - Holochelus lineolatus Reitter, 1902 [by monotypy; ??]
- Holocnemus Brenske, 1894a
  - Holocnemus tomentosus Brenske, 1894a [by monotypy; ??]
- Holomelia Brenske, 1891 [LR Arrow 1944b]
  - Holomelia mirabilisBrenske, 1891 [by monotypy; NZSC]
- Holopycnia Brenske, 1896b
  - Holopycnia celebesiaca Brenske, 1896b [by monotypy; ??]
- Holorhopaea Britton, 1978
  - Holorhopaea sagata Britton, 1978 [by monotypy; SAMA]
- Holotrichia Hope, 1837 [dups?; status Khan & Ghai 1982a; LR Russian spp. Kalinina 1978; see Lachnosterna Phyllophaga]

SUBGENUS Amphitrichia Brenske, 1894a [as genus Arrow 1944b]

- - Holotrichia amboinae Brenske, 1894a [by monotypy; ??]

SUBGENUS Eotrichia Medvedev, 1951 [as genus; Chang 1964; *valid Kalinina 1978; ???]

- - Holotrichia titanis Reitter, 1902

SUBGENUS Eretusa Reitter, 1902

- - Holotrichia nigra (Redtenbacher, 1868) [by monotypy; ??]

                                       [=Ancylonycha nigra Redtenbacher, 1868]

SUBGENUS Holotrichia Hope, 1837

                                    [=Ancyclonycha Dejean, 1833] [Arrow 1944b]
                                    [=Neolepidiota Blackburn, 1890a] [Arrow 1917]
                                    [=Phyllophaga Harris, 1827] [Arrow 1944b]

- - Holotrichia hirticula (Knoch, 1801) [by subseqeunt designation; Glasgow 1916; ZMHB]

                                       [=Melolontha hirticula Knoch, 1801]

- - Holotrichia obscura (Blackburn, 1890) [=Holotrichia serrata Farbricius; by monotypy; SAMA]

                                       [=Neolepidiota obscura Blackburn, 1890]

- - Holotrichia serrata (Fabricius, 1787) [by original designation; BMNH]

                                       [=Melolontha serrata Fabricius, 1787]

- - Holotrichia serrata (Fabricius, 1775) [by subsequent designation; Duponchel 1849; BMNH]

                                       [=Melolontha serrata Fabricius, 1775]

SUBGENUS Noanda Reitter, 1902

- - Holotrichia standfussi Brenske, 1892 [by monotypy; ZMHB]

SUBGENUS Pledina Reitter, 1902 [type sp.?]

                                    [=Hippotrichia Arrow, 1948b] [of Pledina; Chang 1964]
                                    [=Miridiba Reitter, 1902] [of Pledina; Chang 1964]
                                    [=Neodontocnema Arrow, 1948b] [of Pledina; Chang 1964; LR Frey 1970]

- - Holotrichia coromandeliana (Blanchard, 1851) [by original designation; MNHN]

                                       [=Ancylonycha coromandeliana Blanchard, 1851]

- - Holotrichia hainana (Arrow, 1948b) [by original designation; BMNH]

                                       [=Hippotrichia hainana Arrow, 1948b]

- - Holotrichia trichophorus (Fairmaire, 1891) [by monotypy; MNHN]

                                       [=Rhizotrogus trichophorus Fairmaire, 1891]

- Homoeoschiza Kolbe, 1894

                                  [=Euryschiza Brenske, 1898] [Moser 1914a]

- - Homoeoschiza aberrans (Gerstaecker, 1873) [by monotypy; ZMHB]

                                       [=Schizonycha aberrans Gerstaecker, 1873]

- - Homoeoschiza salaama (Brenske, 1898) [by monotypy; ZMHB]

                                       [=Euryschiza salaama Brenske, 1898]

- Hoplochelus Blanchard, 1851 [LR Dewailly 1950]

                                  [=Enariodea Fairmaire, 1903] [Dewailly 1950; syn. of Hoplocnemus?]

- - Hoplochelus rhizotrogoides Blanchard, 1851 [by monotypy; MNHN]
  - Hoplochelus villigera (Fairmaire, 1903) [by monotypy; MNHN]

                                       [=Enarioidea villigera Fairmaire, 1903]

- Hoplocnemus Blanchard, 1851
- Hoplolontha Fairmaire, 1904b
  - Hoplolontha paviei Fairmaire, 1904b [by monotypy; MNHN]
- Hoplosternodes Burgeon, 1946a
  - Hoplosternodes splendens Burgeon, 1946a [by monotypy; ??]
- Hoplosternus Guerin-Meneville, 1838 [Oplosternus]
  - Hoplosternus chinensis (Guerin-Meneville, 1838) [by monotypy; ??]

                                       [=Oplosternus chinensis Guerin-Meneville, 1838]

- Hypochrus Fairmaire, 1891
  - Hypochrus brevicollis Fairmaire, 1891 [by monotypy; MNHN]
- Hypolepida Britton, 1978
  - Hypolepida wilsoni Britton, 1978 [by monotypy; ANIC]
- Hypopholis Erichson, 1847 [LR Machatschke 1955]
  - Hypopholis sommeri Burmeister, 1855 [by subsequent monotypy; MLUH]
- Hypothyce Howden, 1968
  - Hypothyce mixta Howden, 1968 [by monotypy; CNCI]
- Hypotrichia LeConte, 1862
  - Hypotrichia spissipes LeConte, 1862 [by monotypy; MCZC]
- Idiapogonia Arrow, 1916b
  - Idiapogonia laratina Arrow, 1916b [by monotypy; BMNH]
- Idionycha Arrow, 1932
  - Idionycha excisa Arrow, 1932 [by monotypy; BMNH]
- Jalalabadia Balthasar, 1967
  - Jalalabadia povolnyi Balthasar, 1967 [by monotypy; ??]
- Lachnodera Erichson, 1847
  - Lachnodera rufojubata Fairmaire, 1884 [by subsequent monotypy; ZMHB]
- Lachnosterna Hope, 1837 [see Holotrichia Phyllophaga]

SUBGENUS Lachnosterna Hope, 1837

                                    [=Anomalochela Moser] [Arrow 1948]
                                    [=Brahmnina Blanchard, 1851] [Arrow 1948]
                                    [=Holochelus Blanchard, 1851] [Arrow 1948]
                                    [=Holotrichia Hope, 1837] [Arrow 1948; Cartwright & Gordon 1971]

- - Lachnosterna bicolor (Brenske, 1892) [by original designation; ZMHB]

                                       [=Brahmina bicolor Brenske, 1892]

- - Lachnosterna calva (Blanchard, 1851) [by subsequent designation; Arrow 1948; ??]

                                       [=Brahmina calva Blanchard, 1851]

- - Lachnosterna fervida (Fabricius, 1775) [by original designation; OXUM]

                                       [=Melolontha fervida Fabricius, 1775]

- - Lachnosterna serrata (Fabricius, 1787) [by original designation; BMNH]

                                       [=Melolontha serrata Fabricius, 1787]

- - Lachnosterna subseriatus (Reitter, 1889) [by monotypy; ??]

                                       [=Holochelus subseriatus Reitter, 1889]

SUBGENUS Trichestes Erichson, 1848 [Dalla Torre 1912]

- - Lachnosterna pilosicollis (Knoch, 1801) [by subsequent designation; Blanchard 1851; ??]

                                       [=Melolontha pilosicollis Knoch, 1801]

- Lachnota Reitter, 1889
  - Lachnota henningi (Fischer, 1823) [by monotypy; ??]

                                       [=Rhizotrogus henningi Fischer, 1823]

- Lasiexis Semenov-Tian-Shanskii & Medvedev, 1936
  - Lasiexis dilaticollis (Ballion, 1870) [by original designation; ??]

                                       [=Lachnosterna dilaticollis Ballion, 1870]

- Lasiopsis Erichson, 1848

SUBGENUS Brahmina Blanchard, 1851 [*cited Nikolaev 1976; ???] SUBGENUS Lasiobrahmina Medvedev, 1951

- - Lasiopsis duchoni Reitter, 1902 [by original designation; ??]

SUBGENUS Lasiopsis Erichson, 1848 [Medvedev 1951]

- - Lasiopsis henningii (Fischer, 1824) [by subsequent monotypy; Blanchard 1851; ??]

                                       [=Melolontha henningii Fischer, 1824]

- Lasiotropus Reitter, 1913
  - Lasiotropus poonensis Reitter, 1913 [by monotypy; ??]
- Latipalpus Moser, 1921a
  - Latipalpus maxillata (Brenske, 1892) [by original designation; ??]

                                       [=Holotrichia maxillata Brenske, 1892]

- Lecantotrogus Kolbe, 1894
  - Lecantotrogus runsoricus Kolbe, 1894 [by monotypy; ??]
- Lepidiota Kirby, 1828 [LR Australia-Britton 1978]

                                  [=Eulepida Kolbe, 1894] [Arrow 1943]

- - Lepidiota baumanni (Kolbe, 1894) [subsequent designation; Dewailly 1950; ZMHB]

                                       [=Eulepida baumanni Kolbe, 1894]

- - Lepidiota stigma (Fabricius, 1798) [by subsequent designation; Hope 1837; ?BMNH]

                                       [=Melolontha stigma Fabricius, 1798]

- Lepidomela Kolbe, 1894
  - Lepidomela cervina Kolbe, 1894 [by monotypy; ??]
- Lepidotrogus Kolbe, 1894
  - Lepidotrogus conradti Kolbe, 1894 [present designation; ??]
- Lepischiza Moser, 1914a
  - Lepischiza fulvipes (Moser, 1914a) [by original designation; ??]

                                       [=Coniopholis fulvipes Moser, 1914a]

- Leptolepis Ley, 1917 [LR Dewailly 1950]
  - Leptolepis major (Fairmaire, 1903) [by monotypy; ??]

                                       [=Empecta major Fairmaire, 1903]

- Leucopholis Blanchard, 1845 [Dejean 1833?]
  - Leucopholis rorida (Weber, 1801) [present designation; ??]

                                       [=Melolontha rorida Weber, 1801]

- Leucophorus Brenske, 1894a
  - Leucophorus flabellatus Brenske, 1894a [by monotypy; ZMHB]
- Magniscarabaeus Hong, 1988 [date?; et al.]
  - Magniscarabaeus furvus (Hong, 1988) [date?; et al.; by monotypy; fossil-Miocene; ??]

                                       [=Megascarabaeus furvus Hong, 1988]

- Malaisius Arrow, 1941b
  - Malaisius mollis Arrow, 1941b [by monotypy; ??]
- Mascarena Arrow, 1919
  - Mascarena rodriguezi (Waterhouse, 1876) [by monotypy; BMNH]

                                       [=Lachnosterna rodriguezi Waterhouse, 1876]

- Megacoryne Britton, 1987
  - Megacoryne carnabyorum Britton, 1987 [by monotypy; ANIC]
- Meganoxia Reitter, 1902
  - Meganoxia orita Reitter, 1902 [by monotypy; ??]
- Megarhopaea Britton, 1978
  - Megarhopaea gigas (Lea, 1916) [by monotypy; SAMA]

                                       [=Pararhopaea gigas Lea, 1916]

- Megistophylla Burmeister, 1855

                                  [=Hecatomnus Fairmaire, 1891] [Arrow 1944b]

- - Megistophylla grandicornis (Fairmaire, 1891) [by monotypy; MNHN]

                                       [=Hecatomnus grandicornis Fairmaire, 1891]

- - Megistophylla jughuhni Burmeister, 1855 [by monotypy; MLUH]
- Melolontha Fabricius, 1775

SUBGENUS Apropyga Medvedev, 1951

- - Melolontha aceris Faldermann, 1835 [by original designation; ??]

SUBGENUS Melolontha Fabricius, 1775 [Medvedev 1951]

- - Melolontha melolontha (Linne, 1758) [by absolute tautonomy; BMNH]

                                       [=Scarabaeus melolontha Linne, 1758]

SUBGENUS Tocama Reitter, 1902

- - Melolontha rubiginosa Fairmaire, 1889 [by monotypy; MNHN]
- Melolonthoides Dewailly, 1950
  - Melolonthoides rothschildi Dewailly, 1950 [by monotypy; MNHN]
- Metatrogus Britton, 1978
  - Metatrogus septuosus Britton, 1978 [by original designation; ANIC]
- Microlontha Petrovitz, 1967
  - Microlontha aphodioides Petrovitz, 1967 [by monotypy; ??]
- Microphylla Kraatz, 1890
  - Microphylla paupera (Hampe, 1852) [by monotypy; ??]

                                       [=Anoxia paupera Hampe, 1852]

- Microrhopaea Lea, 1920
  - Microrhopaea flavipennis Lea, 1920 [by monotypy; SAMA]
- Microtrichia Brenske, 1900 [LR Arrow 1927;type sp.?]
- Miltotrogus Reitter, 1902 [as subgenus of Rhizotrogus; Baraud 1977; LR Nonvellier 1965]
  - Miltotrogus aequinoctialis (Herbst, 1790) [by subsequent designation; Medvedev 1951; ??]

                                       [=Melolontha aequinoctialis Herbst, 1790]

- Miolachnosterna Wickham, 1914
  - Miolachnosterna tristoides Wickham, 1914 [by monotypy; MCZC]
- Monotropus Erichson, 1847
  - Monotropus nordmanni Blanchard, 1851 [by subsequent monotypy; MNHN]
- Nanorhopaea Britton, 1978
  - Nanorhopaea testacea (Britton, 1978) [by monotypy; SAMA]

                                       [=Nanorrhopaea testacea Britton, 1978]

- Nematophylla Arrow, 1906
  - Nematophylla rugosa Arrow, 1906 [by monotypy; BMNH]
- Ochranoxia Kraatz, 1888 [LR Mevedev 1951]
  - Ochranoxia semiflava (Kraatz, 1883) [by monotypy; ZMHB]

                                       [=Anoxia semiflava Kraatz, 1883]

- Octoplasia Brenske, 1892
  - Octoplasia lineata (Sharp, 1876) [present designation; ??]

                                       [=Lachnosterna lineata Sharp, 1876]

- Oligophylla Kraatz, 1894a
  - Oligophylla detrita (Fairmaire, 1875) [present designation; MNHN]

                                       [=Anoxia detrita Fairmaire, 1875]

- Oreotrogus Kolbe, 1897 [LR Decelle 1973]
  - Oreotrogus volkensi Kolbe, 1897 [by monotypy; BMNH;MNHN;ZMHB]
- Othnonius Olliff, 1890
  - Othnonius batesii Olliff, 1890 [by monotypy; AMSA]
- Pachrodema Blanchard, 1851
  - Pachrodema castanea Blanchard, 1851 [present designation; MNHN]
- Pahangia Arrow, 1948b
  - Pahangia costata Arrow, 1948b [by monotypy; BMNH]
- Panotrogus Reitter, 1902
  - Panotrogus myschenkovi (Ballion, 1871) [by monotypy; ??]

                                       [=Rhizotrogus myschenkovi Ballion, 1871]

- Pararhopaea Blackburn, 1911
  - Pararhopaea callabonnensis (Blackburn, 1894) [by monotypy; SAMA]

                                       [=Rhopaea callabonnensis Blackburn, 1894]

- Pectinichelus Ballion, 1871
  - Pectinichelus rhizotrogoides Ballion, 1871 [by monotypy; ??]
- Pegylis Erichson, 1847

                                  [=Pegylidius Peringuey, 1904] [Arrow 1943]

- - Pegylis mashunus (Peringuey, 1904) [by monotypy; SAMC]

                                       [=Pegylidius mashunus Peringuey, 1904]

- - Pegylis morio (Blanchard, 1851) [by subsequent monotypy; MNHN]

                                       [=Pegylius morio Blanchard, 1851]

- Pentaphylla Dewailly, 1950
  - Pentaphylla gieseckiei Dewailly, 1950 [by monotypy; MNHN]
- Pentelia Brenske, 1891 [LR Arrow 1944b]
  - Pentelia discedens (Sharp, 1881) [by monotypy; ??]

                                       [=Lachnosterna discedens Sharp, 1881]

- Phiara Brenske, 1897
  - Phiara flava Brenske, 1897 [by monotypy; ??]
- Philacelota Heller, 1900
  - Philacelota submaculata Heller, 1900 [present designation; ??]
- Pholidochris Kolbe, 1894
  - Pholidochris dorhni (Quedenfeldt, 1884)

                                       [=Ancylonycha dorhni Quedenfeldt, 1884]

- Phyllophaga Harris, 1827 [see Holotrichia Lachnosterna; dup??]

SUBGENUS Chirodines Bates, 1888

- - Phyllophaga zunilensis (Bates, 1888) [by monotypy; BMNH]

                                       [=Chirodines zunilensis Bates, 1888]

SUBGENUS Chlaenobia Blanchard, 1850 [LR Chapin 1935]

- - Phyllophaga ciliatipes (Blanchard, 1851) [by monotypy; MNHN]

                                       [=Chlaenobia ciliatipes Blanchard, 1851]

SUBGENUS Cnemarachis Saylor, 1942

- - Phyllophaga vandinei (Smyth, 1917) [by original designation; ??; note, jr. synonym of Ancylonycha Arrow 1948]

                                       [=Lachnosterna vandinei Smyth, 1917]

SUBGENUS Eugastra LeConte, 1856

- - Phyllophaga cribrosa (LeConte, 1853) [by subsequent designation; Saylor 1942; MCZC]

                                       [=Tostegoptera cribrosa LeConte, 1853]

SUBGENUS Listrochelus Blanchard, 1851

- - Phyllophaga laportei (Blanchard, 1851) [by monotypy; MNHN]

                                       [=Listrochelus laportei Blanchard, 1851]

SUBGENUS Phyllophaga Harris, 1827

                                    [=Abcrana Saylor, 1942] [as subgenus of Cnemarachis; Sanderson 1951]
                                    [=Ancyclonycha Dejean, 1833] [see Holotrichia]
                                    [=Clemora Saylor, 1942] [Sanderson 1951]
                                    [=Endrosa LeConte, 1856] [??]
                                    [=Gynnis LeConte, 1856] [??]
                                    [=Holotrichia Hope, 1837] [as synonym of Ancyclonycha Saylor 1942]
                                    [=Lachnosterna Hope, 1837] [??]
                                    [=Trichestes Blanchard, 1851] [??]

- - Phyllophaga debilis (LeConte, 1856) [by monotypy; MCZC]

                                       [=Gynnis debilis LeConte, 1856]

- - Phyllophaga fervida (Fabricius, 1775) [by original designation; OXUM]

                                       [=Melolontha fervida Fabricius, 1775]

- - Phyllophaga hirticula (Knoch, 1801) [by subseqeunt designation; Glasgow 1916; ZMHB]

                                       [=Melolontha hirticula Knoch, 1801]

- - Phyllophaga pilosicollis (Knoch, 1801) [by monotypy; ZMHB]

                                       [=Melolontha pilosicollis Knoch, 1801]

- - Phyllophaga quercus (Knoch, 1801) [by monotypy; ZMHB]

                                       [=Melolontha quercus Knoch, 1801]

TYPE SPECIES Phyllophaga serrata (Fabricius, 1775) [DUP; by subsequent designation; Duponchel 1849; BMNH; note see Holotrichia Hope; Arrow 1948 designates Ancylonycha hogardi Blanchard 1851 apecies]

                                       [=Melolontha serrata Fabricius, 1775]

- - Phyllophaga serrata (Fabricius, 1787) [by original designation; BMNH]

                                       [=Melolontha serrata Fabricius, 1787]

- - Phyllophaga smith (Arrow, 1912) [by original designation; BMNH]

                                       [=Phytalus smith Arrow, 1912]

SUBGENUS Phytalus Erichson, 1847 [LR Saylor 1939; Frey 1975a]

- - Phyllophaga pubereus (Mannerheim, 1829) [by subsequent designation; Saylor 1942; ??]

                                       [=Melolontha pubereus Mannerheim, 1829]

SUBGENUS Tostegoptera Blanchard, 1851

- - Phyllophaga lanceolata (Say, 1824?) [by monotypy; ANSP]

                                       [=Melolontha lanceolata Say, 1824?]

SUBGENUS Triodonyx Saylor, 1942

- - Phyllophaga gigantissima (Saylor, 1942) [by original designation; ??]

                                       [=Triodonyx gigantissima Saylor, 1942]

- Plectrodes Horn, 1867

                                  [=Parathyce Hardy, 1974] [new]

- - Plectrodes carpenteri LeConte, 1876 [by origina ldesignation; MCZC]
  - Plectrodes pubescens Horn, 1867 [by monotypy; MCZC]
- Pollaplonyx Waterhouse, 1875a [LR Nomura 1970]

SUBGENUS Bunbunius Nomura, 1970

- - Pollaplonyx reticulata (Murayama, 1941) [by original designation; ??]

                                       [=Holotrichia reticulata Murayama, 1941]

SUBGENUS Polloplonyx Waterhouse, 1875a [Nomura 1970]

- - Pollaplonyx flavidus Waterhouse, 1875a [by monotypy; BMNH]
- Polyenaria Dewailly, 1950
  - Polyenaria ciliata Dewailly, 1950 [by monotypy; MNHN]
- Polylamina Hardy, 1974
  - Polylamina pubescens (Cartwright, 1939) [by monotypy; USNM]

                                       [=Polyphylla pubescens Cartwright, 1939]

- Polyphylla Harris, 1841 [LR Young 1988; Dewailly 1992]

SUBGENUS Centralasiobia Semenov-Tian-Shanskii, 1902 [1901 fide Medvedev 1951]

- - Polyphylla shestakowi Semenov-Tian Shanskii, 1900 [by monotypy; ZMAS]

SUBGENUS Grananoxia Brenske, 1890

- - Polyphylla annamensis (Fleutiaux, 1887) [by monotypy; MNHN]

                                       [=Granida annamensis Fleutiaux, 1887]

SUBGENUS Granida Motschulsky, 186 [as genus; ??]

- - Polyphylla albolineata (Motschulsky, 1861) [by monotypy; ??]

                                       [=Granida albolineata Motschulsky, 1861]

SUBGENUS Gynexophylla Medvedev, 1951

- - Polyphylla laticollis Lewis, 1887 [by original designation; ??]

SUBGENUS Mesopolyphylla Medvedev, 1951

- - Polyphylla irrorata (Gebler, 1841) [by original designation; ZMAS]

                                       [=Melolontha irrorata Gebler, 1841]

SUBGENUS Polyphylla Harris, 1841

- - Polyphylla variolosa (Hentz, 1830) [by subsequent designation; Young 1988; ??]

                                       [=Melolontha variolosa Hentz, 1830]

SUBGENUS Xerasiobia Medvedev, 1951

                                    [=Macronoxia Crotch, 1873] [type sp.?]

- - Polyphylla albus (Pallas, 1773) [by original designation; ??]

                                       [=Scarabaeus albus Pallas, 1773]

- Polyphyllum Blanchard, 1851
  - Polyphyllum rubrescens Blanchard, 1851 [by monotypy; MNHN]
- Proagosternus Blanchard, 1851 [LR Dewailly 1950]
  - Proagosternus niveus Blanchard, 1851 [by subsequent designation; Dewailly 1950; MNHN]
- Proseconius Kolbe, 1894
  - Proseconius capito (Gerstaecker, 1873) [by monotypy; ??]

                                       [=Schizonycha capito Gerstaecker, 1873]

- Proteroschiza Brenske, 1897
  - Proteroschiza wituensis Brenske, 1897 [by monotypy; ZMHB]
- Pseudenaria Fairmaire, 1901 [LR Dewailly 1950]
  - Pseudenaria hexaphylla Fairmaire, 1901 [by monotypy; MNHN]
- Pseudoapterogyna Escalera, 1914 [see de Peyerimhoff 1945-1949; ??; type sp.?]

                                  [=Apterogyna Reiche, 1882] [subgenus of Rhizotrogus Dalle Torre 1912]

- Pseudodiplotaxis Nonfried, 1894
  - Pseudodiplotaxis albosetosa Nonfried, 1894 [by monotypy; ??]
- Pseudoholophylla Blackburn, 1911 [LR Britton 1978]

                                  [=Holophylla Burmeister, 1855] [not Erichson 1847; Blackburn 1911]
                                  [=Paralepidiota Blackburn, 1911] [Britton 1978]

- - Pseudoholophylla furfuracea (Burmeister, 1855) [by monotypy; MLUH]

                                       [=Holophylla furfuracea Burmeister, 1855]

- - Pseudoholophylla lepidoptera (Blackburn, 1912) [by subsequent monotypy; SAMA]

                                       [=Paralepidiota lepidoptera Blackburn, 1912]

- Pseudolontha Fairmaire, 1897b
  - Pseudolontha canaliculata Fairmaire, 1897b [present designation; ??]
- Pseudopantrogus Petrovitz, 1969 [type sp.?]
- Pseudopholis Duvivier, 1892
  - Pseudopholis squamulosa Duvivier, 1892 [by monotypy; ??]
- Pseudosymmachia Dalle Torre, 1912

                                  [=Metabolus Fairmaire, 1887] [Arrow 1944b]
                                  [=Symmachia Brenske, 1892] [not Hubner 1816]

- - Pseudosymmachia chinensis (Brenske, 1892) [by monotypt; MNHN]

                                       [=Symmachia chinensis Brenske, 1892]

- - Pseudosymmachia tumidifrons (Fairmaire, 1887) [by monotypy; MNHN]

                                       [=Metabolus tumidifrons Fairmaire, 1887]

- Pseudotrematodes du Val, 1860
  - Pseudotrematodes frivaldszkyi (Menetries, 1836) [by monotypy; ZMAS]

                                       [=Rhizotrogus frivaldszkyi Menetries, 1836]

- Psilonychus Burmeister, 1855
  - Psilonychus eckloni Burmeister, 1855 [present designation; MLUH]
- Psilopholis Brenske, 1892
  - Psilopholis grandis (Laporte, 1840) [present designation; ??]

                                       [=Tricholepis grandis Laporte, 1840]

- Ramilia Kolbe, 1894a

                                  [=Syntaxopholis Brenske, 1903a] [Burgeon 1946a]

- - Ramilia kuiluensis (Brenske, 1898) [by monotypy; NHRS]

                                       [=Pseudopholis kuiluensis Brenske, 1898]

- - Ramilia pruinosa (Harold, 1878) [by monotypy; ??]

                                       [=Rhopaea pruinosa Harold, 1878]

- Redotus Fairmaire, 1860b
  - Redotus rufulus Fairmaire, 1860b [by monotypy; MNHN]
- Rhabdopholis Burmeister, 1855

                                  [=Hoplobrachium Boheman, 1857] [Dohrn 1863]

- - Rhabdopholis albostriata Burmeister, 1855 [by monotypy; MLUH]
  - Rhabdopholis costipennis (Boheman, 1857) [present designation; NHRS]

                                       [=Hoplobrachium costipennis Boheman, 1857]

- Rhadinolontha Arrow, 1930
  - Rhadinolontha mimetica Arrow, 1930 [by monotypy; BMNH]
- Rhizoproctus Kolbe, 1894a
  - Rhizoproctus aurescens Kolbe, 1894a [by monotypy; ??]
- Rhizotrogus Berthold, 1827 [type sp.?]

SUBGENUS Apterogyna Reiche, 1882 SUBGENUS Rhizocolax Motschulsky, 1860 [as subg. of Brahmina Dalle Torre 1912] SUBGENUS Xanthotrogus Reitter, 1902

- - Rhizotrogus sieversi Reitter, 1896 [by subsequent designation; Medvedev 1951; ??]

SUBGENUS Xesotrogus Reitter, 1902

- - Rhizotrogus validus Kraatz, 1884 [by monotypy; ZMHB]
- Rhopaea Erichson, 1847 [LR Britton 1978]
  - Rhopaea verreauxi Blanchard, 1851 [by subsequent monotypy; Blanchard 1851; MNHN]
- Rhyxicephalus Moser, 1921a
  - Rhyxicephalus rugifrons Moser, 1921a [by monotypy; ZMHB]
- Rhyxiderma Moser, 1924a
  - Rhyxiderma celebensis Moser, 1924a [by monotypy; ZMHB]
- Rubilepis Dewially, 1950
  - Rubilepis camuseti Dewailly, 1950 [by original designation; MNHN]
- Schismatocera des Cottes, 1872
  - Schismatocera nitidula (des Cottes) [by monotypy; ??]

                                       [=Adoretus nitidula des Cottes]

- Schistocometa Brenske, 1902b [see Brahmina]
  - Schistocometa brenskei (Reitter, 1900) [by monotypy; ??]

                                       [=Brahmina brenskei Reitter, 1900]

- Schizonycha Dejean, 1833 [LR Pope1960; Arrow 1944a]

SUBGENUS Etischiza Brenske, 1898 [as genus; Burgeon 1946]

- - Schizonycha heudelotti Blanchard, 1851 [by monotypy; MNHM]

SUBGENUS Schizonycha Dejean, 1833

                                    [=Atys Reiche, 1850] [not de Montfort 1810; Strand 1942]
                                    [=Atysilla Strand, 1942]

- - Schizonycha globator (Fabricius, 1781) [by subsequent designation; Pope 1960; BMNH]

                                       [=Scarabaeus globator Fabricius, 1781]

- - Schizonycha samenensis (Reiche, 1850) [by montypy; BMNH]

                                       [=Atys samenensis Reiche, 1850]

- Schoenherria Burmeister, 1855
  - Schoenherria serrulata) [MLUH; ??]

                                       [=SMelolontha serrulata]

- Sebakue Peringuey, 1904 [LR Decelle 1960]

                                  [=Sebakwe Decelle, 1960] [misspelling ?]

- - Sebakue coriacea Peringuey, 1904 [by monotypy; SAMC]
- Semienaria Dewailly, 1950
  - Semienaria descarpentriesi Dewailly, 1950 [by original designation; MNHN]
- Shangaia Lucas, 1920

                                  [=Holotrochus Brenske, 1894a] [not Erichson]

- - Shangaia vestitus (Brenske, 1894a) [by monotypy; ZMHB]

                                       [=Holotrochus vestitus Brenske, 1894a]

- Sophrops Fairmaire, 1887

SUBGENUS Onychosophrops Frey, 1972

- - Sophrops brevisetosa Frey, 1972 [by monotypy; ??]

SUBGENUS Sophrops Fairmaire, 1887

                                    [=Microtrichia Brenske, 1900b] [Arrow 1946]

- - Sophrops parviceps Fairmaire, 1887 [by monotypy; MNHN]
  - Sophrops puttalama (Brenske, 1900b) [present designation; ZMHB]

                                       [=Microtrichia puttalama Brenske, 1900b]

- Spaniolepis Kolbe, 1894a
  - Spaniolepis excavata Kolbe, 1894a [by monotypy; ZMHB]
- Spathoschiza Arrow, 1902
  - Spathoschiza debilis Arrow, 1902 [by monotypy; BMNH]
- Stenopegylis Arrow, 1943
  - Stenopegylis cylindrica Arrow, 1943 [by monotypy; BMNH]
- Stephanopholis Brenske, 1896c [SEE DECELLE 1968]
  - Stephanopholis philippinensis Brenske, 1896c [present designation; ??]
- Suntemnonycha Peringuey, 1904
  - Suntemnonycha collusor Peringuey, 1904 [by monotypy; SAMC]
- Syngeneschiza Brenske, 1898
  - Syngeneschiza tarsata Brenske, 1898 [by monotypy; ZMHB]
- Terebrogaster Lacroix, 1988/89?
  - Terebrogaster pygidialis (Waterhouse, 1882) [BMNH; ??]

                                       [=Lepidiota pygidialis Waterhouse, 1882]

- Thyce LeConte, 1856 [LR Hardy 1974]
  - Thyce squamicollis LeConte, 1856 [by monotypy; MCZC]
- Toxospathius Deyrolle & Fairmaire, 1878
  - Toxospathius inconstans Fairmaire, 1878 [by montypy; ??]
- Trematodes Falderman, 1835
  - Trematodes tenebrioides (Pallas, 1871) [??]

                                       [=Scarabaeus tenebrioides Pallas, 1871]

- Tricholepis Blanchard, 1851 [LR Dewailly 1950]
  - Tricholepis niveopilosa Blanchard, 1851 [by monotypy; MNHN]
- Trichoschiza Moser, 1917b [type sp.?; ??]
- Trigonocnemus Kraatz, 1894
  - Trigonocnemus hauseri (Kraatz, 1894) [by monotypy; ??]

                                       [=Trigonocnemis hauseri Kraatz, 1894]

- Trinoxia Brenske, 1894a
  - Trinoxia cyphonotoides Brenske, 1894 [by monotypy; ??]
- Unguiculenaris Dewailly, 1950
  - Unguiculenaris unguicularis (Moser, 1919) [by original designation; ZMHB]

                                       [=Encya unguicularis Moser, 1919]

- Vadonaria Dewailly, 1950
  - Vadonaria scultipennis (Fairmaire, 1898) [by original designation; MNHN]

                                       [=Enaria scultipennis Fairmaire, 1898]

- Xenotrogus Britton, 1978
  - Xenotrogus vitiensis (Fairmaire, 1879) [by original designation; MNHM]

                                       [=Rhopaea vitiensis Fairmaire, 1879]

- Zietzia Blackburn, 1894
  - Zietzia geologa Blackburn, 1894 [by monotypy; SAMA]
Tribù Pachydemini Reitter, 1902 [LR neotropical genera Martinez 1975]
- Achloa Erichson, 1840

                                  [=Nanarcta Blanchard, 1851] [Burmeister 1855]

- - Achloa helvola Erichson, 1840 [present designation; ZMHB]
  - Achloa vulpecula (Blanchard, 1851) [by monotypy; MNHN; note, N. vulpecula Blanchard is a jr. synonym of A. helvola Erichson (Burmeister 1855)]

                                       [=Nanarcta vulpecula Blanchard, 1851]

- Acylochilus Ohaus, 1909

                                  [=Acyclochilus Ohaus, 1909] [spelling?]

SUBGENUS Acylochilus Martinez, 1975

- - Acylochilus strumosus Ohaus, 1909 [by subsequent designation; Martinez 1975; ZMHB]

SUBGENUS Acylochoides Martinez, 1975

- - Acylochilus persimilis Martinez, 1959 [by original designation; ??]
- Aglaphyra Brenske, 1896a
  - Aglaphyra setosa Brenske, 1896a [by monotypy; ZMHB]
- Alaia Petrovitz, 1979
  - Alaia sexdentata Petrovitz, 1979 [by monotypy; ??]
- Anahi Martinez, 1958
  - Anahi guaraniticus Martinez, 1958 [by monotypy; ??]
- Anomolyna Fairmaire, 1897
  - Anomolyna picticollis Fairmaire, 1897 [by monotypy; ZMHB]
- Anomonyx Saylor, 1940

                                  [=Anomalonyx Moser, 1921b] [not Weise; Saylor 1940]

- - Anomonyx uruguayensis (Moser, 1921b) [by monotypy; ZMHB]

                                       [=Anomalonyx uruguayensis Moser, 1921b]

- Atanyproctus Petrovitz, 1954
  - Atanyproctus simplicitarsis Petrovitz, 1954 [by monotypy; ??]
- Benedictia Sanderson, 1939 [Pleocomidae?]
  - Benedictia pilosa Sanderson, 1939 [by monotypy; SEMC]
- Brenskiella Berg, 1898

                                  [=Burmeisteria Schickendantz, 1868] [not Gray 1865]
                                  [=Pachychaeta Brenske, 1896a] [not Big.]

- Burmeisteriellus Berg, 1898
  - Burmeisteriellus flavomicans (Brenske, 1896a) [by monotypy; ZMHB]

                                       [=Pachychaeta flavomicans Brenske, 1896a]

- - Burmeisteriellus mirablilis (Schickendantz, 1868) [by monotypy; ??]

                                       [=Burmisteria mirablilis Schickendantz, 1868]

- Campylophyllus Decelle, 1979
  - Campylophyllus somalicus Decelle, 1979 [by monotypy; MZUF]
- Castanochilus Ohaus, 1909
  - Castanochilus bruchianus Ohaus, 1909 [by monotypy; ZMHB]
- Cephaloncheres Brenske, 1898
  - Cephaloncheres ugogoensis Brenske, 1898 [by monotypy; ZMHB]
- Clitopa Erichson, 1847
  - Clitopa bohemani Blanchard, 1851 [by subsequent monotypy; MNHN]
- Cyclomera Klug, 1855

                                  [=Diaclaspus Brenske, 1896a] [new]
                                  [=Paraclitopa Waterhouse, 1875c] [new]
                                  [=Periclitopa Brenske, 1896a] [new]

- - Cyclomera delagoensis (Brenske, 1896a) [by monotypy; ZMHB]

                                       [=Diaclaspus delagoensis Brenske, 1896a]

- - Cyclomera dispar Klug, 1855 [present designation; ZMHB]
  - Cyclomera fischeri (Brenske, 1896a) [ZMHB]

                                       [=Periclitopa fischeri Brenske, 1896a]

- - Cyclomera lanuginosa (Waterhouse, 1875a) [by monotypy; BMNH]

                                       [=Paraclitopa lanuginosa Waterhouse, 1875a]

- Elaphocera Gene, 1836

SUBGENUS Elaphocera Gene, 1836 [Reitter 1902]

- - Elaphocera longitarsis (Illiger, 1803) [by subsequent designation; Medevedev 1952; ??]

                                       [=Melolontha longitarsis Illiger, 1803]

SUBGENUS Elaphocerida Reitter, 1902 [LR Baraud 1979]

- - Elaphocera emarginata (Gyllenhal, 1817) [by subsequent designation; Medvedv 1952; ??]

                                       [=Melolontha emarginata Gyllenhal, 1817]

- Elaphocerella Moser, 1919b
  - Elaphocerella variabilis Moser, 1919b [by monotypy; ZMHB]
- Europtron Marseul, 1867 [Baraud 1977]
  - Europtron gracile Marseul, 1867 [by monotypy; ??]
- Faargia Martinez, 1982
  - Faargia gentilii (Martinez, 1975) [by monotypy; ??]

                                       [=Myloxena gentilii Martinez, 1975]

- Fossocarus Howden, 1961
  - Fossocarus creoleorum Howden, 1961 [by monotypy; FMNH]
- Goniorrhina Quedenfeldt, 1888
  - Goniorrhina flaviceps Quedenfeldt, 1888 [by monotypy; ZMHB]
- Gronocarus Schaeffer, 1927
  - Gronocarus autumnalis Schaeffer, 1927 [by monotypy; ??]
- Hemictenius Reitter, 1897
  - Hemictenius magnitarsus Reitter, 1897 [present designation; ??; not Tanyproctus opacus Ballion 1870 as stated by Medvedev 1952]
- Howdenocarus Hardy, 1978
  - Howdenocarus mexicanus Hardy, 1978 [by monotypy; HAHC]
- Idiochelyna Arrow, 1916a
  - Idiochelyna pectoralis (Walker) [by original designation; BMNH; ??]

                                       [=Isonychus pectoralis Walker]

- Kraseophylla Peringuey, 1904
  - Kraseophylla distincta Peringuey, 1904 [by monotypy; SAMC]
- Kryzhanovskia Nikolaev & Kabakov, 1977
  - Kryzhanovskia olegi Nikolaev & Kabakov, 1977 [by monotypy; ??]
- Leuretra Erichson, 1846 [Blanchard cites 1847]
  - Leuretra ectoralis Erichson, 1846 [by monotypy; ZMHB]
- Lichniops Gutierrez Alonso, 1946
  - Lichniops barberoi Gutierrex Alonso, 1946 [by monotypy; ??]
- Lichniopsoides Martinez, 1953
  - Lichniopsoides breyeri Martinez, 1953 [by monotypy; ??]
- Longicrura Frey, 1974a
  - Longicrura fusca (Frey, 1974a) [by monotypy; ZSMC]

                                       [=Longicrurua fusca Frey, 1974a]

- Luispenaia Martinez, 1972
  - Luispenaia paradoxa Martinez, 1972 [by monotypy; ??]
- Macrophylla Hope, 1837 [see Evans 1988]

                                  [=Eucyclophylla Waterhouse, 1875c] [new]
                                  [=Pleiophylla Peringuey, 1904] [new]

- - Macrophylla lata (Waterhouse, 1875c)

                                       [=Eucyclophylla lata Waterhouse, 1875c]

- - Macrophylla longicornis (Fabricius, 1787) [by original designation; BMNH]

                                       [=Melolontha longicornis Fabricius, 1787]

- - Macrophylla vestita (Peringuey, 1904) [by monotypy; SAMC]

                                       [=Pleiophylla vestita Peringuey, 1904]

- Madiniella Chalumeau & Gruner, 1976
  - Madiniella christinae Chalumeau & Gruner, 1976 [by monotypy; ZSMC]
- Myloxena Ohaus, 1909
  - Myloxena vestita Ohaus, 1909 [by monotypy; ??]
- Myloxenoides Martinez, 1975
  - Myloxenoides tandilensis Martinez, 1975 [by monotypy; AMIC]
- Neogutierrezia Martinez, 1953 [LR Martinez 1974]
  - Neogutierrezia mirablis Martinez, 1953 [by monotypy; ??]
- Oedanomerus Waterhouse, 1875c
  - Oedanomerus hirsutus Waterhouse, 1875c [by monotypy; BMNH]
- Onochaeta Erichson, 1847
  - Onochaeta porcata (Swartz, 1817) [by monotypy; NHRS]

                                       [=Melolontha porcata Swartz, 1817]

- Onochirus Kolbe, 1891
  - Onochirus fulvescens Kolbe, 1891 [by monotypy; ZMHB]
- Ootoma Blanchard, 1850 [syn. of Pachydema?; see Baraud 1985]
  - Ootoma bipartita (Brulle, 1838) [present designation; ??]

                                       [=Pachydema bipartita Brulle, 1838]

- Otoclinus Brenske, 1896a
  - Otoclinus gracilipes Brenske, 1896a [by monotypy; ??]
- Pachnessa Brenske, 1894b
  - Pachnessa nicobarica (Redtenbacher, 1868) [by monotypy; ??]

                                       [=Pachydema nicobarica Redtenbacher, 1868]

- Pachycolus Erichson, 1847
  - Pachycolus madagascarensis Blanchard, 1851 [by subsequent monotypy; MNHN]
- Pachydema Laporte, 1832b [LR Baraud 1985]

SUBGENUS Pachydema Laporte, 1832b [Reitter 1902]

- - Pachydema nigricans Laporte, 1832 [junior synonym of Melolontha hirticollis Fabricus 1787; by monotypy; ??]

SUBGENUS Phygotoxeuma Brenske, 1897

- - Pachydema theriyi Brenske, 1897

SUBGENUS Sparophysa Brumeister, 1855 [type sp.?]

- Pachydemocera Reitter, 1902
  - Pachydemocera lucidicollis (Kraatz, 1882) [by present designation; ZMHB]

                                       [=Elaphocera lucidicollis Kraatz, 1882]

- Parapetiia Martinez, 1961
  - Parapetiia camba Martinez, 1961 [by monotypy; ??]
- Pasaphylla Peringuey, 1904
  - Pasaphylla libens Peringuey, 1904 [by monotypy; SAMS]
- Paulosawaya Martinez & D'Andretta, 1955
  - Paulosawaya ornatissima Martinez & D'Andretta, 1955 [by monotypy; ??]
- Pentacoryna Moser, 1926
  - Pentacoryna bruchi Moser, 1926 [by monotypy; ZMHB]
- Periproctus Kolbe, 1910
  - Periproctus orestius Kolbe, 1910 [by monotypy; ZMHB]
- Perissosoma Waterhouse, 1875c
  - Perissosoma aenescens Waterhouse, 1875c [by monotypy; BMNH]
- Peritryssus Reitter, 1918
  - Peritryssus excisus Reitter, 1918 [by monotypy; ??]
- Phalangonyx Reitter, 1889 [see Decelle 1982]
  - Phalangonyx coniceps Reitter, 1889 [by ??; ??]
- Phalangosoma Quedenfeldt, 1884
  - Phalangosoma mechowi Quedenfeldt, 1884 [by monotypy; ??]
- Phobetus Leconte, 1856 [LR Hardy 1978]
  - Phobetus comatus LeConte, 1856 [by monotypy; MCZC]
- Pimelomera Moser, 1924b
  - Pimelomera schwetzi Moser, 1924b [by monotypy; ZMHB]
- Pleistophylla Peringuey, 1904
  - Pleistophylla singularis Peringuey, 1904 [by monotypy; SAMC]
- Protoclitopa Decelle, 1979
  - Protoclitopa tetraphylla Decelle, 1979 [by monotypy; BMNH]
- Protoctenius Medvedev, 1952
  - Protoctenius bogatshevii Medevedev, 1952 [by monotypy; ??]
- Pseudachloa Peringuey, 1904
  - Pseudachloa leonina Peringuey, 1904 [by monotypy; SAMC]
- Pseudoleuretra Martinez & D'Andretta, 1956
  - Pseudoleuretra bokermanni Martinez & D'Andretta, 1956 [by monotypy; ??]
- Pseudoliogenys Moser, 1919b
  - Pseudoliogenys flavidus Moser, 1919 [by monotypy; ZMHB]
- Pseudopachydema Balthasar, 1930
  - Pseudopachydema caucasica Balthasar, 1930 [by monotypy; ??]
- Pseustophylla Peringuey, 1904

                                  [=Ramoutsa Peringuey, 1908] [new]

- - Pseustophylla fervida (Boheman, 1857) [by monotypy; NHRS]

                                       [=Clitopa fervida Boheman, 1857]

- - Pseustophylla pretoriana Peringuey, 1904 [by monotypy; SAMC]
- Ptyophis Redtenbacher, 1868
  - Ptyophis paulseni (Philippi, 1864) [by monotypy; ??]

                                       [=Tetraphyllus paulseni Philippi, 1864]

- Scapanoclypeus Evans, 1987b
  - Scapanoclypeus aberrans (Frey, 1974) [by original designation; ZSMC]

                                       [=Trichinopus aberrans Frey, 1974]

- Scaphorhina Quedenfeldt, 1884

                                  [=Aipeiopsis Peringuey, 1904] [new]

- - Scaphorhina crinipes Quedenfeldt, 1884 [by monotypy; MNHN]
  - Scaphorhina hirsuta (Peringuey, 1904) [subsequent designation; Evans 1987c; SAMC]

                                       [=Aipeiopsis hirsuta Peringuey, 1904]

- Sebaris Laporte, 1840
  - Sebaris palpalis Laporte, 1840 [by monotypy; ??]
- Selomothus Fairmaire, 1891
  - Selomothus diversiceps Fairmaire, 1891 [by monotypy; MNHM not located]
- Sparrmannia Laporte, 1840 [LR Evans 1989]

                                  [=Cephalotrichia Hope, 1837] [new]
                                  [=Leocaeta Dejean, 1833] [new]
                                  [=Leontochaeta Erichson, 1847] [new]

- - Sparrmannia alopex (Fabricius, 1787) [by original designation; ??]

                                       [=Melontha alopex Fabricius, 1787]

- - Sparrmannia alopex (Fabricius, 1787) [present designation; ??]

                                       [=Melolontha alopex Fabricius, 1787]

- - Sparrmannia alopex (Fabricius, 1787) [by subsequent designation; Evans 1989; ??]

                                       [=Melolontha alopex Fabricius, 1787]

- - Sparrmannia alopex (Fabricius, 1787) [by monotypy; ??]

                                       [=Melolontha alopex Fabricius, 1787]

- Synclitopa Kolbe, 1898
  - Synclitopa simulatrix Kolbe, 1898 [by monotypy; ZMUH]
- Tanyproctoides Petrovitz, 1971
  - Tanyproctoides arabicus Petrovitz, 1971 [by monotypy; ??]
- Tanyproctus Faldermann, 1835 [LR Baraud 1979]

SUBGENUS Brachydema [see Petrovitz 1963; ??] SUBGENUS Tanyproctocera Reitter, 1902

- - Tanyproctus festivus Burmeister, 1855 [by subsequent designation; Medvedev 1952; MLUH]

SUBGENUS Tanyproctus Faldermann, 1835 [Reitter 1902]

- - Tanyproctus persicus Menetries, 1832 [by subsequent designation; Medevedev 1952; ??]

SUBGENUS Tetraproctus Khnzoryan, 1953

- - Tanyproctus antennatus Khnzoryan, 1953 [by monotypy; ??]
- Taphrocephala Quedenfeldt, 1888
  - Taphrocephala polita Quedenfeldt, 1888 [by monotypy; MHNP]
- Thoracotrichia Decelle, 1968a
  - Thoracotrichia somalica Decelle, 1968a [by monotypy; MRAC]
- Tlaocera Peringuey, 1904
  - Tlaocera saga Peringeuy, 1904 [by monotypy; SAMC]
- Tosevskiana Pavicevic, 1985
  - Tosevskiana inexpectata (Pavicevic, 1985) [by monotypy; ??]

                                       [=Tosevkiana inexpectata Pavicevic, 1985]

- Trichinopus Waterhouse, 1874c

                                  [=Damara Peringuey, 1904] [new]
                                  [=Deuterohapalopus Dalle Torre, 1912] [pro Hapalopus; new]
                                  [=Hapalopus Brenske, 1903] [not Ausserer]

- - Trichinopus chuni (Brenske, 1903) [by monotypy; ZMHB]

                                       [=Hapalopus chuni Brenske, 1903]

- - Trichinopus flavipennis Waterhouse, 1874c [BMNH]
  - Trichinopus villosula (Peringuey, 1908) [by monotypy; SAMC]

                                       [=Damara villosula Peringuey, 1908]

- Trichiodera Burmeister, 1855
  - Trichiodera bicarinata (Gyllenhal, 1817) [by monotypy; ??]

                                       [=Melolontha bicarinata Gyllenhal, 1817]

- Triodontonyx Moser, 1921a
  - Triodontonyx ukerewensis Moser, 1921a [by monotypy; ??]
Tribù Pachytrichini Burmeister, 1855
- Pachytricha Hope, 1841 [LR Allsopp 1990]

                                  [=Pachytrichia Burmeister, 1855] [incorrect subsequent spelling]

- - Pachytricha castanea Hope, 1841 [by monotypy; BMNH]
Tribù Phyllotocidiini Britton, 1957
- Phyllotocidium Blackburn, 1898 [LR Britton 1957]
  - Phyllotocidium macleayi (Blackburn, 1891) [by monotypy; BMNH]

                                       [=Cheiragra macleayi Blackburn, 1891]

Tribù Scitalini Britton, 1957 [key to genera Britton 1987]
- Byrromorpha Blackburn, 1892 [LR Britton 1987]
  - Byrromorpha verres Blackburn, 1892 [present designation; SAMA]
- Colpochelyne Britton, 1987
  - Colpochelyne dux Britton, 1987 [by monotypy; ANIC]
- Colpochilodes Blackburn, 1898 [LR Britton 1987]
  - Colpochilodes raucipennis Blackburn, 1898 [by monotypy; SAMA]
- Gnaphalopoda Reiche, 1860 [LR Britton 1987]

                                  [=Xylostygnus Broun, 1886] [Given 1960; LR Given 1952]

- - Gnaphalopoda piceus (Broun, 1881) [by monotypy; BMNH]

                                       [=Xylostygnus piceus Broun, 1881]

- - Gnaphalopoda varians Reiche, 1860 [by monotypy; ??; cited as Rhisotrogus leptopoda Montrouzier 1860 fide Britton 1992]
- Hadrops Britton, 1987
  - Hadrops halei Britton, 1987 [by original designation; SAMA]
- Homolotropus Macleay, 1871 [LR Britton 1970]

                                  [=Homalotropus Dalla Torre, 1912] [subsequent incorrect spelling]

- - Homolotropus luridipennis Macleay, 1871 [by monotypy; AMSA]
- Idanastes Britton, 1987
  - Idanastes abditus Britton, 1987 [by original designation; QMBA]
- Ophropyx Britton, 1987
  - Ophropyx hispida (Blackburn, 1898) [by original designation; BMNH]

                                       [=Frenchella hispida Blackburn, 1898]

- Phorine Britton, 1987
  - Phorine anomala (Blackburn, 1907) [by original designation; BMNH]

                                       [=Byrromorpha anomala Blackburn, 1907]

- Protoleura Britton, 1987
  - Protoleura nana Britton, 1987 [by original designation; ANIC]
- Scitala Erichson, 1842 [LR Britton 1987]
  - Scitala sericans Erichson, 1842 [by monotypy; ZMHB]
- Sericesthis Boisduval, 1835 [LR Britton 1987]

                                  [=Anodontonyx Sharp, 1890] [Britton 1987]

- - Sericesthis geminata Boisduval, 1835 [by subsequent designation; Blackburn 1907; unk.]
  - Sericesthis vigilans (Sharp, 1890) [by subsequent designation; Britton 1987]

                                       [=Anodontonyx vigilans Sharp, 1890]

- Synchilus Britton, 1955a
  - Synchilus gisleni Britton, 1955a [by monotypy; MZLU]
- Telura Erichson, 1842 [LR Britton 1987]
  - Telura vitticollis Erichson, 1842 [by monotypy; ZMHB]
- Xyrine Britton, 1987
  - Xyrine carnei Britton, 1987 [by original designation; ANIC]
- Xyroa Britton, 1987
  - Xyroa polita Britton, 1987 [by original designation; ANIC]
- Xyrodes Britton, 1987
  - Xyrodes calorata (Blackburn, 1907) [by original designation; BMNH]

                                       [=Frenchella calorata Blackburn, 1907]

Tribù Sericini Dalla Torre, 1912 [subfamily?]

SUBTRIBE Phyllotocina Machatschke, 1959 [LR Britton 1957]

                                  [=Macrothops MacLeay, 1864] [preoccupied; Blackburn 1898]

- Ancylonyx Britton, 1957
  - Ancylonyx vittatus (MacLeay, 1887) [by original designation; MAMU]

                                       [=Phyllotocus vittatus MacLeay, 1887]

- Anthotocus Britton, 1957
  - Anthotocus lurida (MacLeay, 1864) [by original designation; MAMU]

                                       [=Cheiraga lurida MacLeay, 1864]

- Cheiragra Macleay, 1864 [LR Britton 1957]
  - Cheiragra pusillus (Blanchard, 1851) [by subsequent designation; Britton 1957; MNHN?]

                                       [=Phyllotocus pusillus Blanchard, 1851]

- Cheirodontus Britton, 1957
  - Cheirodontus basicollis (Lea, 1919) [by original designation; QMBA]

                                       [=Phyllotocus basicollis Lea, 1919]

- Cheirora Britton, 1957
  - Cheirora interstitialis (Blackburn, 1898) [by subsequent monotypy; BMNH]

                                       [=Cheirramphica interstitialis Blackburn, 1898]

- Neophyllotocus Blackburn, 1898 [LR Britton 1957]
- Opistocus Britton, 1957
  - Opistocus obscurus (Lea, 1920) [by monotypy; SAMA]

                                       [=Phyllotocus obscurus Lea, 1920]

- Phyllotocus Fischer, 1823 [LR Britton 1957]

                                  [=Macrothops Boisduval, 1835] [Britton 1957]
                                  [=Macrotops Castlenau, 1840] [subsequent misspelling]

- - Phyllotocus macleayi Fischer, 1823 [by monotypy; UMBB]
  - Phyllotocus praeusta (Boisduval, 1835) [present designation; unknown fide Britton 1992]

                                       [=Macrothops praeusta Boisduval, 1835]

- - Phyllotocus rostrata (Macleay, 1864) [by original designation; MAMU]

                                       [=Macrothops rostrata Macleay, 1864]

- Sphaeroscelis Burmeister, 1855 [LR Britton 1957]

                                  [=Cheirramphica Blackburn, 1898] [Britton 1957]

- - Sphaeroscelis pectoralis Burmeister, 1855 [by monotypy; MLUH]
  - Sphaeroscelis pubescens (Blackburn, 1898) [present designation; BMNH]

                                       [=Cheirramphica pubescens Blackburn, 1898]

SUBTRIBE Sericina Brenske, 1897 [LR genera of western Palearctic Baraud 1965; genera of South America Frey 1973]

- Anomalophylla Reitter, 1887 [Reitter 1902 for subgenus]

                                  [=Melaserica Brenske, 1898] [Reitter 1902]
                                  [=Xorema Reitter, 1902] [as subgenus of Anomalophylla; Nikolajev ??]

- - Anomalophylla moupinea (Fairmaire, 1891) [by present designation; MNHN]

                                       [=Serica moupinea Fairmaire, 1891]

- - Anomalophylla thibetana (Brenske, 1898) [by monotypy; ZMHB]

                                       [=Melaserica thibetana Brenske, 1898]

- - Anomalophylla tristicula Reitter, 1887 [by monotypy; ??]
- Anomioserica Arrow, 1947 [publ. date 1946 appeared Jan. 1947]
  - Anomioserica flavipes Arrow, 1946 [by monotypy; BMNH]
- Aphaenoserica Brenske, 1901 [emendation by Dalle Torre in Schenkling 1912 p.45, 46]

                                  [=Aphenoserica Brenske, 1900:77]

- - Aphaenoserica fallax (Brenske, 1901) [by monotypy; ZMHB]

                                       [=Aphenoserica fallax Brenske, 1901]

- Apotriodonta Baraud, 1964
  - Apotriodonta hispanica (Baraud, 1962) [by monotypy; MNMS]

                                       [=Triodonta hispanica Baraud, 1962]

- Archoserica Brenske, 1901
  - Archoserica bogosana Brenske, 1901 [by monotypy; ZMHB]
- Astaena Erichson, 1847b [LR Frey 1972]
  - Astaena tridentata Erichson, 1847b [by??; ZHMB]
- Athlia Erichson, 1835 [LR Martinez 1974]

SUBGENUS Athlia Erichson, 1835

                                    [=Rivera Germain, 1903] [Saylor 1946]

- - Athlia plebeja (Burmeister, 1855) [by monotypy; MLUH]

                                       [=Rivera plebeja Burmeister, 1855]

- - Athlia rustica Erichson, 1835 [by monotypy; ZMHB]

SUBGENUS Dihymenonyx Gutierrez, 1949 [Martinez 1974]

- - Athlia herrerai (Gutierrez Alonso, 1949) [by monotypy; ??]

                                       [=Dihymenonyx herrerai Gutierrez Alonso, 1949]

SUBGENUS Sayloria Frey, 1973

- - Athlia bicoloripes (Saylor)

                                       [=Astanea bicoloripes Saylor]

- Aulacoserica Brenske, 1900a [LR west African spp. Frey 1968]
  - Aulacoserica flava Brenske, 1902 [present designation; ZMHB]
- Balbera Fairmaire, 1898
  - Balbera reflexa (Fairmaire, 1897) [by monotypy; MNHN]

                                       [=Blabera reflexa Fairmaire, 1897]

- Bielliana Baraud, 1980
  - Bielliana abysinnica (Brenske, 1902) [by original designation; ZMHB]

                                       [=Holoschiza abysinnica Brenske, 1902]

- Blepharotoma Blanchard, 1850 [LR Frey 1973]
  - Blepharotoma tarsalis (Blanchard, 1850) [by monotypy; MNHN]

                                       [=Blepharatoma tarsalis Blanchard, 1850]

- Calloserica Brenske, 1894a
  - Calloserica tigrina Brenske, 1894a [by monotypy; ZMHB]
- Camentoserica Brenske, 1901
  - Camentoserica livida (Boheman, 1860) [by monotypy; NHRS]

                                       [=Serica livida Boheman, 1860]

- Chaetoserica Brenske, 1897a
  - Chaetoserica cymosa Brenske, 1897a [by monotypy; ZMHU]
- Charioserica Brenske, 1900a
  - Charioserica striata Brenske, 1900a [by monotypy; ZMHB]
- Chrysoserica Brenske, 1897a
  - Chrysoserica auricoma Brenske, 1897a [by present designation; ZMHB]
- Clypeoserica Frey, 1971
  - Clypeoserica producta Frey, 1971 [by monotypy; MZHF]
- Comaserica Brenske, 1897a
  - Comaserica dapsilis Brenske, 1897a [present designation; ZMHB]
- Conioserica Brenske, 1901
  - Conioserica eisermanni Brenske, 1901 [by monotypy; ZMHB]
- Coronoserica Brenske, 1902
  - Coronoserica beata Brenske, 1902 [by monotypy; ZMHB]
- Corynoserica Moser, 1918
  - Corynoserica tetraphylla Moser, 1918 [by monotypy; ZMHB]
- Cyphoserica Brenske, 1900a
  - Cyphoserica mukengeana Brenske, 1900a [by monotypy; ZMHB]
- Deroserica Moser, 1915
  - Deroserica pulchra Moser, 1915 [present designation; ZMHB]
- Diaphoroserica Peringuey, 1904

SUBGENUS Diaphoroserica Peringuey, 1904

- - Diaphoroserica mashona (Peringuey, 1904) [by monotypy; SAMC]

                                       [=Diaphoserica mashona Peringuey, 1904]

SUBGENUS Dissotoxus Fairmaire, 1899

- - Diaphoroserica insignicornis (Fairmaire, 1899) [by monotypy; MNHM]

                                       [=Dissotoxus insignicornis Fairmaire, 1899]

- Doleroserica Peringuey, 1904
  - Doleroserica gentilis Peringeuy, 1904 [present designation; SAMC]
- Doxocalia Brenske, 1901
  - Doxocalia superba Brenske, 1901 [by monotypy; ZMHB]
- Emphania Erichson, 1847

                                  [=Heptomera Blanchard, 1850] [Burmeister 1855]

- - Emphania metallica (Blanchard, 1850) [by monotypy; MNHN]

                                       [=Heptomera metallica Blanchard, 1850]

- - Emphania metallica (Blanchard, 1850) [by subsequent monotypy; Burmeister 1855; MNHN]

                                       [=Heptomera metallica Blanchard, 1850]

- Eriphoserica Brenske, 1900a
  - Eriphoserica camentoides Brenske, 1900a [by monotypy; ZMHB]
- Etiserica Peringuey, 1904
  - Etiserica simplex Peringuey, 1904 [by monotypy; SAMC]
- Euphoresia Brenske, 1900a
  - Euphoresia butneri Brenske, 1900a [present designation; ZMHB]
- Euserica Reitter, 1896 [LR Baruad 1965a]

SUBGENUS Euserica Reitter, 1902

- - Euserica mutata (Gyllenhal, 1817) [by ?]

                                       [=Melolontha mutata Gyllenhal, 1817]

SUBGENUS Eusericula Reitter, 1902

- - Euserica modesta (Fairmaire, 1881) [by monotypy; MNHN]

                                       [=Serica modesta Fairmaire, 1881]

- Gastroserica Brenske, 1898
  - Gastroserica sulcata Brenske, 1898 [present designation; ZMHB]
- Glaphyserica Brenske, 1900a
  - Glaphyserica humeralis Brenske, 1900a [present designation; ZMHB]
- Glycyserica Brenske, 1900a
  - Glycyserica depravata Brenske, 1900a [by monotypy; ZMHB]
- Gynaecoserica Brenske, 1898
  - Gynaecoserica pellecta (Brenske, 1896) [by monotypy; ZMHB]

                                       [=Serica pellecta Brenske, 1896]

- Hellaserica Baraud & Nicolas, 1966
  - Hellaserica elongata (Reitter, 1887) [by original designation; ??]

                                       [=Homaloplia elongata Reitter, 1887]

- Hemiserica Brenske, 1894a
  - Hemiserica nasuta Brenske, 1894a [by monotypy; ZMHB]
- Heteroserica Brenske, 1900a
  - Heteroserica paradoxa Brenske, 1900a [by monotypy; ZMHB]
- Homaloplia Stephens, 1830 [emendation Erichson 1847]

                                  [=Omaloplia Stephens]

SUBGENUS Acarina Baraud, 1965b

- - Homaloplia spiraeae (Pallas, 1776) [by original designation; MNHN]

                                       [=Scarabaeus spiraeae Pallas, 1776]

SUBGENUS Homaloplia Baraud, 1965b

- - Homaloplia ruricola (Fabricius, 1775) [by ?]

                                       [=Melolontha ruricola Fabricius, 1775]

- Homaloserica Brenske, 1901
  - Homaloserica fessa Brenske, 1901 [by monotypy; ZMHB]
- Hyboserica Peringuey, 1904
  - Hyboserica globuliformis (Brenske, 1892) [present designation; ZMHB]

                                       [=Serica globuliformis Brenske, 1892]

- Hymenochelus Reitter, 1890
  - Hymenochelus distincta (Uhagon, 1876) [by monotyopy; ??]

                                       [=Serica distincta Uhagon, 1876]

- Hymenoplia Eschscholtz, 1830
- Hyposerica Brenske, 1897a
  - Hyposerica definitiva Brenske, 1897a [present designation; ZMHB]
- Lamproserica Brenske, 1901
  - Lamproserica mombasana Brenske, 1901 [present designation; ZMHB]
- Lepidoserica Nikolaev, 1979
  - Lepidoserica polyphylla (Moser) [by original designation; ZMHB; ??]

                                       [=Pachyserica polyphylla Moser]

- Leucoserica Reitter, 1896

                                  [=Brachyserica Brenske, 1897] [?Dalle Torre 1912]

- - Leucoserica arenicola (Solsky, 1876) [??]

                                       [=Serica arenicola Solsky, 1876]

- - Leucoserica arenicola (Solsky, 1876) [by original designation; ??]

                                       [=Serica arenicola Solsky, 1876]

- Leuroserica Arrow, 1947 [publ. date 1946 appeared Jan. 1947]
  - Leuroserica fulgida Arrow, 1946 [by original designation; BMNH]
- Loricatarsa Frey, 1969 [type sp. ?]
- Maladera Mulsant & Rey, 1871 [LR Baraud 1965b; type sp.?]

SUBGENUS Eumaladera Nomura, 1967

- - Maladera nitididorsis Nomura, 1967 [by original designation; ??]

SUBGENUS Maladera Lewis, 1895 [type sp.?]

                                    [=Amaladera Reitter, 1896] [with Maladera Aserica Nikolaev 1977]
                                    [=Aserica Lewis, 1895] [Arrow 1927; with Autoserica Chapin 1932; with Maladera Pope 1961]
                                    [=Autoserica Brenske, 1898a] [Arrow 1927; Pope 1961]
                                    [=Cycloserica Reitter, 1896] [with Maladera Nikolaev 1977 in subgenus Aserica]
                                    [=Lepiserica Brenske, 1901] [Burgeon 1942]
                                    [=Macroserica Medvedev, 1952] [subg.; with Maladera Nikolaev 1977 in subgenus Aserica]
                                    [=Neoserica Brenske, 1894a] [as subgenus of Serica; genus Brenske 1898; Arrow 1946; Pope 1961]
                                    [=Omalodera Reitter, 1896] [as subgenus of Amaladera; ??]

- - Maladera cavifrons (Reitter, 1896) [by original designation; ??]

                                       [=Amaladera cavifrons Reitter, 1896]

- - Maladera euphorbiae (Burmeister, 1855) [by original designation; MLUH]

                                       [=Serica euphorbiae Burmeister, 1855]

- - Maladera excisiceps (Reitter, 1896) [by monotypy; ??]

                                       [=Cycloserica excisiceps Reitter, 1896]

- - Maladera japonica (Motschulsky, 1857) [by original designation; ZMUM; ZMHB; Arrow (1927) designated Serica secreta Brenske 1897 based on Lewis' misidentification of S. japonica. This case must be decided by the Zoological Commission; see Art. 67i; 70b.]

                                       [=Serica japonica Motschulsky, 1857]

- - Maladera moreli (Coquerel, 1866) [present designation; ??]

                                       [=Serica moreli Coquerel, 1866]

- - Maladera piceorufa (Fairmaire, 1888) [by subsequent designation; Arrow 1927; MNHN; ??]

                                       [=Serica piceorufa Fairmaire, 1888]

- - Maladera punctatissima (Falderman, 1835) [by original designation; ??]

                                       [=Serica punctatissima Falderman, 1835]

- - Maladera suturata (Brenske, 1894a) [by monotypy; ??]

                                       [=Serica suturata Brenske, 1894a]

- - Maladera ukamina (Brenske, 1901) [present designation; ZMHB]

                                       [=Lepiserica ukamina Brenske, 1901]

SUBGENUS Paramaladera Nikolaev, 1979

- - Maladera alternata (LeConte, 1856) [by original designation; MCZC]

                                       [=Serica alternata LeConte, 1856]

- Manoserica Moser, 1924
  - Manoserica disparicornis (Quedenfeldt, 1888) [by original designation; ZMHB]

                                       [=Serica disparicornis Quedenfeldt, 1888]

- Mericserica Brenske, 1897a
  - Mericserica oberthuri (Brenske, 1898) [by subsequent monotypy; ZMHB]

                                       [=Meriserica oberthuri Brenske, 1898]

- Mesoserica Brenske, 1901

SUBGENUS Mesoserica Brenske, 1901 [Matsumura 1914]

- - Mesoserica sachalinensis (Matsumura, 1914)

                                       [=Sericania sachalinensis Matsumura, 1914]

- - Mesoserica transvaalensis Brenske, 1901 [by monotypy; ZMHB]

SUBGENUS Sericania Motschulsky, 1860 [Matsumura 1911; type sp.?]

- Nedymoserica Brenske, 1901
  - Nedymoserica flavida Brenske, 1901 [by monotypy; ZMHB]
- Nematoserica Arrow, 1917
  - Nematoserica coerulea Arrow, 1917 [by monotypy; BMNH]
- Neomaladera Baraud, 1965a
  - Neomaladera barbara (Lucas, 1846) [by original designation; MNHN]

                                       [=Homaloplia barbara Lucas, 1846]

- Nepaloserica Frey, 1965a
  - Nepaloserica procera Frey, 1965a [by monotypy; ZSMC]
- Neuroserica Brenske, 1901
  - Neuroserica fulvescens (Blanchard, 1851) [by monotypy; MNHN]

                                       [=Emphania fulvescens Blanchard, 1851]

- Octotemna Blanchard, 1850
  - Octotemna singularis Blanchard, 1850 [by monotypy; MNHN]
- Onychoserica Moser, 1916
  - Onychoserica flabellata Moser, 1916 [by present designation; ??]
- Orthoserica Brenske, 1901 [preoccupied not Warren 1896 in Lep. Geom.]
  - Orthoserica fulvastra Brenske, 1902 [by subsequent monotypy; ZMHB]
- Ovoserica Frey, 1968
  - Ovoserica testaceipennis Frey, 1968 [by monotypy; ??]
- Oxyserica Brenske, 1900a
  - Oxyserica pygidialis Brenske, 1900a
- Pachyderoserica Moser, 1920
  - Pachyderoserica crassicollis Moser, 1920 [by monotypy; ZMHB]
- Pachypoides Fairmaire, 1884
  - Pachypoides limbipennis Fairmaire, 1884 [by monotypy; MNHN]
- Pachyserica Brenske, 1897a [?1898]
  - Pachyserica rubrobasalis Brenske, 1897a [by monotypy; ZMHB]
- Paragynaecoserica Khan & Ghai, 1982b
  - Paragynaecoserica pubescens Khan & Ghai, 1982b [by monotypy; ??]
- Paraserica Reitter, 1896
  - Paraserica griesea (Motschulsky, 1866) [by original designation; ??]

                                       [=Serica griesea Motschulsky, 1866]

- Paratriodonta Baraud, 1964
  - Paratriodonta morio (Fabricius, 1792) [by original designation; BMNH]

                                       [=Melolontha morio Fabricius, 1792]

- Parthenoserica Brenske, 1900a
  - Parthenoserica sulcata Brenske, 1900a
- Philoserica Brenske, 1901
  - Philoserica vittata (Blanchard, 1851) [present designation; MNHN]

                                       [=Brachyphylla vittata Blanchard, 1851]

- Phylloserica Brenske, 1900a
  - Phylloserica unicolor (Vollenhoven, 1869) [present designation; ZMHB]

                                       [=Pleophylla unicolor Vollenhoven, 1869]

- Pleophylla Erichson, 1847
  - Pleophylla fasciatipennis Blanchard, 1851 [present designation; MLUH]
- Plesioserica Brenske, 1900a [emendation Dalle Torre 1912]

                                  [=Plaesioserica Brenske]

- - Plesioserica viridula (Brenske, 1900a) [by monotypy; ZMHB]

                                       [=Plaesioserica viridula Brenske, 1900a]

- Plotopuserica Brenske, 1900a
  - Plotopuserica darwiniania Brenske, 1900a [by monotypy; ZMHB]
- Plusioserica Brenske, 1900a
  - Plusioserica virescens (Kunckel, 1887) [by monotypy; ZMHB]

                                       [=Omaloplia virescens Kunckel, 1887]

- Podoserica Breit, 1912
  - Podoserica reitteri Breit, 1912 [by monotypy; ??]
- Psednoserica Brenske, 1900a
  - Psednoserica amoena Brenske, 1900a [by monotypy; ZMHB]
- Pseudomaladera Nikolaev, 1982
  - Pseudomaladera koltzei (Reitter, 1897) [by original designation; ??]

                                       [=Serica koltzei Reitter, 1897]

- Pseudosericania Kobayashi, 1980
  - Pseudosericania gibiventris Kobayashi, 1980 [by monotypy; ??]
- Raysymmela Saylor, 1947 [LR Frey 1973]
  - Raysymmela bruchi (Moser, 1924) [by original designation; ZMHB]

                                       [=Symmela bruchi Moser, 1924]

- Rhyncoserica Burgeon, 1942
  - Rhyncoserica rostrata Burgeon, 1942 [by monotypy; MRAC?]
- Rhyncosymmela Frey, 1974
  - Rhyncosymmela pallida Frey, 1974 [by monotypy; ??]
- Selaserica Brenske, 1897a
  - Selaserica nitida (Candeze, 1861) [by monotypy; ISNB]

                                       [=Serica nitida Candeze, 1861]

- Serica Macleay, 1819

                                  [=Lasioserica Brenske, 1897a] [Arrow 1946a]
                                  [=Ophthalamoserica Brenske, 1897a] [subg.; Arrow 1946a; see Trichoserica]

- - Serica brunnea (Linnaeus, 1758) [by ??; BMNH; see Arrow 1946a]

                                       [=Scarabaeus brunnea Linnaeus, 1758]

- - Serica calva (Brenske, 1898a) [present designation; ZMHB]

                                       [=Lasioserica calva Brenske, 1898a]

- Somatoserica Brenske, 1899b
  - Somatoserica sikorae Brenske, 1899b [by monotypy; ZMHB]
- Sphecoserica Brenske, 1899b
  - Sphecoserica gracilipennis Brenske, 1899b [by monotypy; ZMHB]
- Stenoserica Brenske, 1901
  - Stenoserica kolbei Brenske, 1901 [present designation; ZMHB]
- Straliga Fairmaire, 1901
  - Straliga croceicollis (Fairmaire, 1901) [by monotypy; MNHN]

                                       [=Straglia croceicollis Fairmaire, 1901]

- Symmela Erichson, 1835
  - Symmela instabilis Erichson, 1835 [present designation; ZMHB]
- Taiwanoserica [?; type sp.?]
- Tamnoserica Brenske, 1900a
  - Tamnoserica mutans (Brenske, 1900a) [present designation; ZMHB]

                                       [=Emphania mutans Brenske, 1900a]

- Tephraeoserica Brenske, 1901
  - Tephraeoserica hauseri Brenske, 1901 [by monotypy; ZMHB]
- Teraserica Brenske, 1897a
  - Teraserica timorana Brenske, 1897a [by monotypy; ZMHB]
- Thrymoserica Brenske, 1901
  - Thrymoserica fabulosa Brenske, 1901 [by monotypy; ZMHB]
- Trachyserica Brenske, 1900a
  - Trachyserica longitarsa Bresnke, 1900a [by monotypy; ZMHB]
- Trichoserica Reitter, 1896

                                  [=Opthalmoserica Brenske, 1897a] [Nomura 1972; see Serica]

- - Trichoserica fulvopubens Reitter, 1896 [by monotypy; ???]
- Triodontella Reitter, 1919 [LR Baraud 1964]

                                  [=Euronycha Peringuey, 1904] [Arrow1917]
                                  [=Hetamius Fairmaire, 1893] [de Peyrerimhoff 1945]
                                  [=Triodonta Mulsant, 1842] [preocc.]

- - Triodontella aquila (Laporte, 1840) [by ??; MNHN]

                                       [=Serica aquila Laporte, 1840]

- - Triodontella demaisoni (Fairmaire, 1893) [by monotypy; MNHN]

                                       [=Hetamius demaisoni Fairmaire, 1893]

- - Triodontella sericans (Fahraeus, 1857) [present designation; NHRS]

                                       [=Triodonta sericans Fahraeus, 1857]

- Trioserica Moser, 1922
  - Trioserica sparsequamosa Moser, 1922 [present designation; ZMHB]
- Trochaloschema Reitter, 1896
  - Trochaloschema iris (Semenow, 1893) [by ??; ??]

                                       [=Serica iris Semenow, 1893]

SUBTRIBE Trochalina Brenske, 1897a

- Ablaberoides Blanchard, 1850
  - Ablaberoides aeneus Blanchard, 1850 [by monotypy; MNHN]
- Antitrochalus Brenske, 1900a
  - Antitrochalus abyssinicus Brenske, 1902 [by subsequent monotypy; ZMHB]
- Bilga Fairmaire, 1893
  - Bilga pictipennis Fairmaire, 1893 [by monotypy; ??]
- Campytrochalus Brenske, 1900
  - Campytrochalus glabriclypealis (Brenske, 1902) [by subsequent monotypy; ZMHB]

                                       [=Campylotrochalus glabriclypealis Brenske, 1902]

- Cephaloserica Brenske, 1900
  - Cephaloserica phthisica Brenske, 1902 [by subsequent monotypy; ZMHB; C. phthisica is a jr. syn. of Autoserica thompsoni (Brenske) 1894 fide Arrow 1946]
- Cyrtotrochalus Brenske, 1900
  - Cyrtotrochalus opacus (Brenske, 1902) [by subsequent monotypy; ZMHB]

                                       [=Cytotrochalus opacus Brenske, 1902]

- Dolerotarsa Peringuey, 1904
  - Dolerotarsa emendatrix Peringuey, 1904 [by monotypy; SAMC]
- Eotrochalus Burgeon, 1943
  - Eotrochalus puncticeps Burgeon, 1943 [present designation; MRAC?]
- Heterotrochalus Moser, 1918
  - Heterotrochalus macrophyllus Moser, 1918 [by monotypy; ZMHB]
- Holoschiza Lansberge, 1886
  - Holoschiza dentilabris Lansberge [by monotypy; ??]
- Loboserica Kolbe, 1914
  - Loboserica gracilis Kolbe, 1914 [by subsequent designation; Zool. Rec. 1968; ZMHB]
- Mesotrochalus Kolbe, 1914
  - Mesotrochalus docilis Kolbe, 1914 [by subsequent designation; Zool. Rec. 1968; ZMHB]
- Microserica Brenske, 1894b [as subgenus of Serica; Brenske 1898]
  - Microserica striola (Brenske, 1894b) [ZMHB; ??]

                                       [=Serica striola Brenske, 1894b]

- Microsericaria Nikolaev, 1980

TYPE SPECIES Microsericaria quadrimaculata (Brenske, 1896) [by monotypy; ??; note Serica quadrimaculata is thpecies of Microserica; by subsequent designation Arrow 1946]

                                       [=Serica quadrimaculata Brenske, 1896]

- Microtrochalus Brenske, 1900
  - Microtrochalus bipunctatus Brenske, 1902 [by subsequent monotypy - ZMHB & is junior synonym of Trochalus plagiger Peringuey 1892; SAMC; Peringuey1904]
- Periserica Brenske, 1898 [LR Arrow 1927]
  - Periserica picta Brenske, 1898 [by monotypy; ZMHB]
- Phyllotrochalus Brenske, 1900
  - Phyllotrochalus montanus Brenske, 1902 [by subsequent monotypy; ZMHB]
- Pseudotrochalus Quedenfeldt, 1884 [type sp. ?]
- Sphaerotrochalus Brenske, 1900
  - Sphaerotrochalus bohmi (Quedenfeldt, 1888) [by subsequent monotypy; Brenske 1902; ??]

                                       [=Pseudotrochalus bohmi Quedenfeldt, 1888]

- Stilbotrochalus Kolbe, 1914
  - Stilbotrochalus nitens Kolbe, 1914 [by monotypy; ZMHB]
- Triacmoserica Kolbe, 1914
  - Triacmoserica stegmanni Kolbe, 1914 [by monotypy; ZMHB]
- Trochaloserica Brenske, 1900a
  - Trochaloserica festiva (Brenske, 1902) [by subsequent monotypy; ZMHB]

                                       [=Trochaloschema festiva Brenske, 1902]

- Trochalus Laporte, 1832

SUBGENUS Epitrochalus Frey, 1960 [as genus; LR Frey 1968]

- - Trochalus zumpti Frey, 1960 [by original designation; ??]

SUBGENUS Trochalus Laporte, 1832 [Frey 1968]

- - Trochalus rotundatus Laporte, 1832 [by ?monotypy; ??; see Laporte 1840]
- Xenotrochalus Moser, 1917a
  - Xenotrochalus mirabilis Moser, 1917a [by monotypy; ZMHB]
Tribù Sericoidini Burmeister, 1855
- Aplodema Blanchard, 1850

                                  [=Haplodema Gemminger & Harold, 1869] [unjustified emendation]

- - Aplodema magellanica (Hombres-Firmas & Jacquelin du Val, 1842) [present designation; MNHN]

                                       [=Brachyphylla magellanica Hombres-Firmas & Jacquelin du Val, 1842]

- Sericoides Guerin-Meneville, 1839

SUBGENUS Accia Curtis, 1845

- - Sericoides lucida (Curtis, 1845) [by monotypy; ??]

                                       [=Accia lucida Curtis, 1845]

SUBGENUS Listronyx Guerin-Meneville, 1839

- - Sericoides nigriceps (Guerin-Meneville, 1839) [by monotypy; ??; a junior synonym of Melolontha testacea Fabricius 1775 in the Sericoides Brenske 1906]

                                       [=Listronyx nigriceps Guerin-Meneville, 1839]

SUBGENUS Paralistronyx Brenske, 1906 [as genus; Martinez 1959]

                                    [=Apterodema Fairmaire, 1884a] [Martinez 1972]
                                    [=Paramaypa Brenske, 1906] [of Paralistronyx Martinez 1959]

- - Sericoides acuticollis (Fairmaire, 1884a) [by monotypy; MNHN]

                                       [=Apterodema acuticollis Fairmaire, 1884a]

- - Sericoides livida (Germain, 1862) [by monotypy; ??]

                                       [=Listronyx livida Germain, 1862]

- - Sericoides orsornoana (Brenske, 1906) [by monotypy; ZMHB]

                                       [=Maypa orsornoana Brenske, 1906]

SUBGENUS Sericoides Guerin-Meneville, 1839

                                    [=Apterodema Fairmaire, 1883] [Martinez 1972]
                                    [=Macrosoma Hope, 1837] [preoccupied]
                                    [=Maypa Blanchard, 1850]

- - Sericoides acuticollis (Fairmaire) [by monotypy; MNHN]

                                       [=Apterodema acuticollis Fairmaire]

- - Sericoides castanea Guerin-Meneville, 1839 [by monotypy; ??]
  - Sericoides chlorosticta (Blanchard, 1850) [by monotypy; MNHN]

                                       [=Maypa chlorosticta Blanchard, 1850]

- - Sericoides glacialis (Fabricius, 1775) [by monotypy; BMNH]

                                       [=Melolontha glacialis Fabricius, 1775]

- Ulata Saylor, 1945b
  - Ulata argentina Saylor, 1945b [CASC]
- Zaburina Saylor, 1945a
  - Zaburina colombiana Saylor, 1945a [by monotypy; CASC]
Tribù Systellopini Sharp, 1877 [LR Allsopp 1989]
- Chilodiplus Sharp, 1877
  - Chilodiplus albertisi Sharp, 1877 [by monotypy; MNHN]
- Enamillus Sharp, 1877
  - Enamillus striatus Sharp, 1877 [by monotypy; MNHN]
- Prochelyna Erichson, 1847

                                  [=Atholerus Sharp, 1877] [Allsopp 1989]
                                  [=Tosotarsus Sharp, 1877] [Allsopp 1989]
                                  [=Toxotarsus Nonfried, 1892] [misspelling of Tosotarsus]

- - Prochelyna heterodoxa Burmeister, 1855 [by subsequent monotypy; Burmeister 1855; SMTD]
  - Prochelyna obscurus (Sharp, 1877) [by monotypy; MNHN]

                                       [=Atholerus obscurus Sharp, 1877]

- - Prochelyna velutinus (Sharp, 1877) [by monotypy; MNHN]

                                       [=Tosotarsus velutinus Sharp, 1877]

- Sarothermus Blackburn, 1907
  - Sarothermus sharpi (Blackburn, 1905) [by original designation; BMNH]

                                       [=Enamillus sharpi Blackburn, 1905]

- Sphyrocallus Sharp, 1877
  - Sphyrocallus brunneus Sharp, 1877 [by monotypy; MNHN]
- Systellopus Sharp, 1877
  - Systellopus obtusus Sharp, 1877 [by subsequent designation; Allsopp 1989; MNHN]
- Trichelasmus Sharp, 1877 [Allsopp 1989]
  - Trichelasmus pilicollis Sharp, 1877 [by monotypy; MNHN]
Tribù Xylonychini Britton, 1957 [key to genera Britton 1979]
- Chariochilus Brenske, 1898

                                  [=Lepidostethaspis Arrow, 1924] [Britton 1979]

- - Chariochilus metallescens Brenske, 1898 [by monotypy; ZMHU]
  - Chariochilus squamosus (Lea, 1919) [by monotypy; SAMA]

                                       [=Stethaspis squamosus Lea, 1919]

- Colymbomorpha Blanchard, 1850 [LR Britton 1957]
  - Colymbomorpha lineata Blanchard, 1850 [by monotypy; MNHN]
- Exostethus Britton, 1979
  - Exostethus brunneus (Britton, 1979) [by monotypy; ANIC]

                                       [=Exostehtus brunneus Britton, 1979]

- Modialis Fairmaire & Germain, 1860
  - Modialis prasinella Fairmaire & Germain, 1860 [by monotypy; ??]
- Mycernus Broun, 1904 [LR Given 1952]
  - Mycernus elegans Broun, 1904 [by monotypy; BMNH]
- Nitorellus Britton, 1957
  - Nitorellus splendida (Lea, 1919) [by monotypy; SAMA]

                                       [=Colymbomorpha splendida Lea, 1919]

- Phyllococerus Waterhouse, 1876
  - Phyllococerus purpurascens Waterhouse, 1876 [by monotypy; BMNH]
- Phytholaema Blanchard, 1850 [LR Saylor 1937]
  - Phytholaema mutabilis Blanchard, 1850 [by monotypy; ??]
- Psilodontria Broun, 1895 [LR Given 1952]
  - Psilodontria viridescens Broun, 1895 [by monotypy; BMNH]
- Pyronota Boisduval, 1835 [LR Given 1952]

                                  [=Calonota Hope, 1837] [Given 1952]

- - Pyronota festiva (Fabricius, 1792) [by original designation; BMNH; see Given 1952]

                                       [=Melolontha festiva Fabricius, 1792]

- - Pyronota festiva (Fabricius, 1792) [by subsequent designation; Given 1952; ??]

                                       [=Melolontha festiva Fabricius, 1792]

- Stethaspis Hope, 1837 [Pope 1982; Watt 1982; LR Given 1952] [=Chlorochiton Arrow, 1903] [??] [=Neosthaspsis Dalle Torre, 1912] [for Stethaspis in error] [=Stethaspis Broun, 1893] [not Hope 1837; not Burmeister 1855; Arrow 1903; Dalle Torre 1912]
  - Stethaspis suturalis (Boisduval, 1835) [by subsequent designation; Arrow 1903; BMNH] [=Melolontha suturalis Boisduval, 1835]
  - Stethaspis suturalis (Boisduval, 1835) [Opinion ???; BMNH] [=Melolontha suturalis Boisduval, 1835]
- Xylonichus Boisduval, 1835 [LR Britton 1957] [=Xylonychus Blanchard, 1851] [incorrect subsequent spelling]
  - Xylonichus eucalypti Boisduval, 1835 [by monotypy; unknown]

^ Family Scarabaeidae (TXT), in Synopsis of the described Coleoptera of the World, Texas A&M University - Department of Entomology. URL consultato il 3 giugno 2014.

[1] Family Scarabaeidae (TXT), in Synopsis of the described Coleoptera of the World, Texas A&M University - Department of Entomology. URL consultato il 3 giugno 2014.

[1]

Utente:Esculapio/Sandbox/Melolonthinae

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